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SUSITNA HYDROELECTRIC PROJECT
.~\PHASE II PFIOGRESS REPORT
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BIG GAME STUDIES
Volume IX . BELUKHA WHALE
Donald G. Calkins
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425 ~ ALASKA DEPARTMENT OF FISH AND GAME
\~;: Submitted to the Alaska Power Authority
o.42e A.prll 1983
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SUSITNA HYDROELECTRIC PROJECT
PHASE II
1982 ANNUAL REPORT
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BIG GAME STUDIES
VOL-. IX BELUKHA WHALE
by Donald G. Calkins
ALASKA DEPARTMENT OF FISH AND GAME
Submi tted to the
Alaska Power Authori ty
Apri 1, . igg~
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ARLIS
Alaska Resources
Library & Information ServiceS
Anchorage, Alaska
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PREFACE
In early 1980,the Alaska Department of Fish and Game contracted
with the Alaska Power Authority to collect information useful in
assessing the impacts of the proposed Susitna Hydroelectric
Proj ect on moose,caribou,wolf,wolverine,black bear,brown
bear and Dall sheep.
The studies were broken into phases which conformed to the
anticipated licensing schedule.,Phase I studies,January 1,1980
to June 30,1982,were intended to _provide information needed to
support a FERC license application.This included general
studies of wildlife populations to determine how each species
used the area and identify potential impact mechanisms.Phase II
studies continued to provide additional information during the
anticipated 2 to 3 year period between application and final FERC
approval of the license.Belukha whales were added to the
species being studied.During Phase I I,we are narrowing the
focus of our studies to evaluate specific impact mechani sms,
quantify impacts and evaluate mi tigation measures.
This is the first annual report of ongoing Phase II studies.In
some cases,objectives of Phase I were continued to provide a
more complete data base.Therefore,this report is not intended
as a complete assessment of the impacts of the Susi tna Hydro-
electric Proj ect on the selected wildlife species.
The information and conclusions contained in these reports are
incomplete and preliminary in nature and subject to change with
further study.Therefore,information contained in these reports
is not to be quoted or used in any publication without the
wri tten permission of the authors..
The reports are organized into the following 9 volumes:
Volume I.
Volume I I.
Volume III.
Volume IV.
Volume V.
Volume VI.
Volume VII.
Volume VIII.
LVolume IX.
Big Game Summary Report
Moose -Downstream
Moose -Upstream
Caribou
Wolf
Black Bear and Brown Bear
Wolverine
Dall Sheep
Belukha Whale
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Summary
Be1ukha surveys were flown in Upper Cook Inlet between May 17 and Aug.
27,1982.A concentration·area was identified nearshore from the
mouth of the Little Susitna River to the mouth of the Beluga River.
Exact timing of use of the area has not been determined,however,the
concentration appeared to build up_in late May and lasted through mid-
June.It is probable that this concentration was in part associated
with calving and breeding although no calves were positively identi-
fied because of generally poor viewing conditions.The concentration
appeared to involve'200 to 300 animals,however accurate counts were
not possible because of.again,poor viewing conditions..The Belukha
concentration near the mouth of the Susitna River appeared to coincide
with the arrival of large numbers of eulachon which spawned in the
lower Susitna River in late May and early June.This run of eulachon
was estimated to total several million fish.King salmon are probably
not particularly important to this concentration of belukhas although
large male belukhas probably do take some king salmon.Probably the
only other salmon species from the Susitna River system available in
sufficient numbers to be considered significant prey to the belukhas
concentrated in late May and early June would be the sockeye.No
information is presently available which would allow conclusions on
belukha predation on salmon smolts from the Susitna River.
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Given the present state of our knowledge,we cannot accurately predict
impacts on Cook Inlet belukhas from the proposed dams on the Susitna
River.It is possible that the overall population could suffer reduc-
tion in numbers both directly by alterations in the habitat,particu-
larly the concentration area near the mouth of the Susitna River and
indirectly by reduction of available food species.
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Introduction
The belukha whale (Defphinapterus Leucas)is a small.toothed whale.
which inhabits arctic and sub-arctic waters.As adults,belukhas are
ail white in coloration,range from 3.1 to 4.4 m in length and weigh
480-1200 kg.Males generally are larger than females.Newborn calves
are a dark gray color.shaded with blue or brown.This coloration is
gradually lost as the animal approaches the age of maturity.
Belukhas range throughout Cook Inlet.concentrating in the upper Inlet
in the spring and summer.and moving to the lower Inlet during the
winter.There is some evidence which suggests that during winters of
heavy ice in Cook Inlet Some of the belukhas may'leave the Inle~
entirely and move across the north Gulf of Alaska to as far away as
Yakutat Bay (Calkins 1979).In the north Gulf of Alaska,belukhas
have been sighted in Shelikof Strait.near Kodiak Island,in Prince
William Sound.and in Yakutat Bay (Fiscus.Braham and Mercer 1976 j
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Harrison and Hall 1978 j Calkins and Pitcher 1978 j Calkins 1979;and
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,~Calkins.unpub~data).There is some evidence which indicates that
some parts of these areas formerly had large numbers of belukha,at
least seasonally.but now the population appears reduced (Calkins,
unpub.data).
The Cook Inlet stock of belukha whales was estimated at 300 to 400
animals by Klinkhart (1966).Recent surveys in the Inlet have shown
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that the population exceeds 400anirnals (Calkins unpub.data).How-
I ever,all surveys of belukhas in Cook Inlet have consisted of aerial
counts of shoreline areas.No complete systematic census of Cook
Inlet belukhas has been completed,therefore no accurate estimate of
the stock can be made.The best information we presently have is that
there are over 400 whales which inhabit Cook Inlet in the summer.
Belukhas are known to feed on a broad assortment of fishes and inver-
etically isolated from other belukha whale stocks.The next nearest
Some evidence exists which suggests that the Cook Inlet stock is gen-
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In EschoJtz Bay belukhas ate
polycheatas and octopus (Seaman et.al 1982).
areas.However the Cook Inlet sample was too small to conclude that
concentrated on rainbow smelt in May and shifted to downstream migrat-
craniological morphology has actually changed in this stock.
ing salmon smolt by June 1.By late June the °Bristol Bay belukhas
utilized by belukhas in Bristol Bay were flounders,lamprey blennys,
prey selection bybelukhas in Cook Inlet.In Bristol Bay belukhas
a small series of skulls from Cook Inlet and compared them to other
were concentrating on adult salmon (Brooks 1954).Other food species
differentiation has taken place in Cook Inlet.He was able to examine
these two stocks.Fay (Pers.Comm)suggests that some morphological
safron cod,sculpins and small amounts of shrimp,isopods,snails,
shrimp and sculpins (Brooks 1954).
stock of belukhas is the population which inhabits Bristol Bay.We
tebrates in other.areas,however no direct information is available on
know of no instance where any interchange has taken place between
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Very little information is available on the belukhas ability to toler-
ate perterbations in it's environment.We know nearly nothing about
the consequence of reducing the food supply or changing the heat
budget of the river,however slight these changes might be.We do
know that belukhas will abandon areas if the environmental perterba-
tions are great enough as was the case in the St.Lawerence River
where belukhas quit using the Manicougan and Outardes Rivers after
they were damned for hydroelectric purposes ..Also human disturbance
can have extraordinary effect as was the case when belukhas failed to
return to the Beluga River in Cook Inlet after heavy hunting pressure.
Methods
Shoreline aerial surveys of upper Cook Inlet (Fig.I)were conducted
periodically from May 17 through August 27,1982 using single engine
aircraft with water landing capability.The surveys were generally
flown at an altitude of 50 to 75 meters and approximately 500'meters
offshore from the local tide water line.When groups of belukhas were
sighted the altitude was increased to 200 meters and the groups were
circled while two observers counted all belukhas sighted.
Turbid water conditions along with short surface times for the beluk-
has prevented accurate and complete counts of all belukhas present.
The numbers of belukhas Gounted represents a minimum number of animals
which were present at the time of the count.An attempt was made to
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Figure 1.Upper Cook Inle.I.'ukha atudr .,.a.
o 'gmlNORTH~..?9
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identify calves in all groups observed by circling and observing at
lower altitude.
Results and Discussion
Belukha surveys are summarized in Table 1.A total of 9 surveys were
flown throughout the period.Each survey covered the same general
area of upper Cook Inlet shoreline,including all areas north of a
line between the north Foreland and moose point.On the June 18th and
June 22nd surveys the survey area was extended to the East and West
Forelands.No neonates were positively identified on any of these
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surveys due to their turbid water conditions.However,on both the
May 17 and the June 4 surveys very dark,small belukhas were sighted.
These could have been newborn calves although this was not determined
because newborn calves and yearlings only differ in length by approxi-
mately 30 cm (John Burns pers.corom.).Determining 30 cm qifference
between animals from an aircraft at 100 to 200 m altitude and moving
at an airspeed of approximately 80 kts with the belukhas in highly
turbid water proved to be an i~possible task.
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T~ble 1.Belukha sureys of upper Cook Inlet May 17,1982 through
Aug 27,1982.
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I~i!l\·May 17
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June 4
June 11
June 18
June 22
July 2
July 8
Aug.5
Aug.27
Number
Sighted
15
10
15
150-200
100
15
20
200-300
108
39
50-75
78
30
15
4
50-75
40
25
46
7
7
63
62
30
21
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Location
Sighted
South of Little Susitna R.
North of Beluga R.
W.pt.Fire Island
Between Susitna R.and Lewis R.
Lewis R.to Beluga R.
SW side Chickaloon Bay
Chickaloon R.Mouth
Lewis R.to Beluga R.
Susitna R.Mouth
Ivan R.'to Beluga R.
Beluga R.and Mouth
Beluga R.to Tyonek village
Chickaloon Bay
Boulder pt.(N.of Kenai)
Mouth of Susitna R.'
Lewis to Theodore R.
Beluga R.and Mouth
McArthur R.and Mouth
Susitna R.to Beluga R.
Beluga R.area
Susitna R.Mouth to Beluga R.
McArthur R.area
Chickaloon Bay
Potter Marsh
Mouth of Beluga R.
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Early in the surveys one area was identified as a concentration area.
This area was just offshore from the water line and extended from the
mouth of the Little Susitna River to just south of the mouth of the
Beluga River.The exact timing of use of this .area has not been
determined.A large number of belukhas could have arrived and
remained in this area prior to th2 beginning of surveys on May 17th:
The pilot of the aircraft for the first survey reported seeing "large
numbers of belugas"near the mouth of the Susitna River in early May.
This concentration lasted through the third week in June as indicated
by the June 18th survey shown in Table 1.However,by the June 22nd
survey we began to see belukhas at other locations in Cook Inlet and
on June 24th,several hundred belukhas were reported in Turnagain Arm
near McHugh Creek and across to Gull Point.
There are several possible explanations which are immediately apparent
for belukhas concentrating in the area near the mouths of the rivers
in the northwest part of Cook Inlet.concentrations in areas of river
mouths in the spring is common to many belukha whale populations.
Seargent and Brodie (1975)considered the primary reason for whales
gathering at this time of year was calving and breeding while Fraker
et.a1.1978 felt that taking advantage of warmer estuarine temper-
atures in·the spring was important to all segments of a belukha popu-
lation,not just the reproductive age classes._They hypothesized that
the thermal advantage to all classes was the primary reason for spring
estuarine aggregations of belukhas,and secondarily,the concentration
areas,may afford some shelter from storms.Fraker et.al (1978)felt
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information was obtained.the total number of fish was estimated in
the several millions.Brooks (1954)fo~d that smelt (Osmerus dentax)
were .important in the diet of belukhas in Bristol Bay very early in
that availability of an important food source was not a major factor
for belukhas concentrating in the Mackenzie estuary because most
whales examined from the native whale harvest had empty stomachs.
However this may be a significant difference from the Cook Inlet stock
of belukhas as well as the Bristol Bay stock.We know from Brooks
(1954)and Lensink (1961)that belukhas do feed in the estuaries in
the spring in Bristol bay and may be primarily attracted to the area
by both downstream migrating smolts and returning adults of several
species of salm0rbs well as an early run of smelt.
Although no detailed abundanceAnadramousFishInvestigations).
Belukhas feeding in estuaries in Cook Inlet in the spring has not been
positively determined.However.the presence and timing of several
species of anadramous fish.similar to the conditions in Bristol Bay.
suggest this to be the case.Probably the single most important fish
species to the belukhas in Cook Inlet in the concentration area in the
spring is the eulachon (Thaleicthys pacificus)which arrives in the
Susitna estuary and enters the river for spawning in two major runs.
The first was detected in the river when sampling began on May 16 and
lasted until approximately May 30 (ADF&G 1982 Adult Anadramous Fish
Investigations).The second run of eulachon entered the river and
covered a nine day period lasting until June 9.The second was con-
sidered to be 4.5 times greater than the first (ADF&G 1982 Adult
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the spring,shortly,after breakup.Smelt are similar in size and
habits to eulachon and probably eulachon are comparable in the diet of
Cook Inlet belukhas.
After the smelt run decreased by the end of May,belukhas in Bristol
Bay switched to dow~stream migrating salmon smolt (Brooks 1954).
Lensink (1961)felt that as the smolt moved out of the estuary and
into Bristol Bay,they apparently ,scattered and became much less vul-
nerable to predation.Thus of six belukhas taken between June 6 and
June 15,none had eaten smolt.If this is also the case in Cook
Inlet,then it is possible that salmon smolt from the Susitna River
may not be an important food source.However,Cook Inlet is somewhat
different from Bristol Bay and particularly at low tide extensive sand
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bars are exposed with the majority of the Inlets water confined to
channels and waterways,considerably smaller then Kvichakor Nushagak
Bays in Bristol Bay.Although the belukhas apparently seldom enter
the Susitna River,the salmon smolt may be concentrated enough at low
tide outside the river mouth to allow the belukhas to feed on them.
Certainly,no realistic estimate of belukha use of salmon smol t iIi.
Cook Inlet can be made without examining stomach contents of the
helukhas.
Adult king salmon (Oncorhynchus tshawytscha)are available .in the
area of the SusitnaRiver (ADF&G,unpub.data),from mid April through
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August with a peak in June.However their value to belukhas as a food
source is questionable.Brooks (1954)did not find adult king salmon
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to be a significant component of the diet of belukhas in Bristol Bay
and speculated that only the smaller adult king salmon were taken by
the largest adult male belukhas.If this is the case in Cook Inlet
also,.probably adult king salmon are taken only in small numbers.
All four of the other species of North American salmon enter the
Susitna River to spawn but probably only the sockeye (Oncorhynchus
nerka)would likely be present in numbers sufficient to provide a sig-
nificant food source during the time period when the belukhas are con-
centrated near the mouth of the Susitna River.However even sockeyes
do not become abundant in upper Cook Inlet until after July 1,after
the concentration of belukhas has dispersed .
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Potential Impacts
Quantification of impacts of the Susitna hydroelectric project on
belukhas at the present time is not possible.This type of proj ect
has the potential for reducing the numbers concentrating near the
mouth of the river by reducing the available food or by altering the
heat budget of the river.However the overall effect on the avail-
ability of anadromous fish to belukhas is predicted to be small.
There may be no alteration of the heat budget of the river realized by
the belukhas at the mouth of the river although very little data are
available to prove this.
Approximately 5 to 8%of the tptal adult salmon returning to the
Susitna River system spawn in the area from Talkeetna to Devil Canyon;
the area which is predicted to be the most heavily impacted by'dam
construction.The slough habitat in this area is predicted to be com-
pletely lost,thereby eliminating approximately 5%of the chum salmon
from the system as well as a small number of sockeye.This means that
a small amount of food in the form of adult churn and sockeye will no
longer be available to the belukhas after dam construction.Since we
have no quantitative measure of the importance of these species to the
belukhas,no estimate of impact can be made except to guess that it
will probably be slight.
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Impacts on the eulachon runs which enter the Susitna River are assumed
to be slight as they remain in the lower reaches of the river (Bruce
Barrett ADF&G pers.comm.).This species may be extremely important
to the belukhas and it is pass ible that any reduction of eulachon
could severly impact the belukhas.
Although most.impacts from either heat budget alteration or food
reduction are likely to be slight,we cannot accurately predict the
overall effect on the belukhas.If a..."'ly environmental perterbations
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effect the belukhas in upper Cook Inlet,it is likely these effects
will take the form of a reduction in the population in Cook Inlet.
Given our pn<sent state of knowledge,a reduction in the belukha popu-
lation of upper Cook Inlet would not be detectable unless it were
greater than a 50%to 75%reduction in the entire population.Even a
reduction of this magnitude could go unnoticed for several years as no
systematic monitoring of the population is planned.
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Recommendations for future.studies
The most immediate information needs for the Cook Inlet belukha popu
lation with resp'ect to the Susitna hydroelectric project is a real
istic population estimate. Generation of such an estimate would
require development of a systematic aerial census of the belukhas in
the entire Inlet from. which a statistically sound estimate could then
be made. Beyond that, future studies should involve, collections of
skulls in order to make an absolute determination on the taxonomic
status of this ~opulation; food habits studies to positively identify
and quantify the importance of food species; and movement studies to
define the geographical range and seasonal movements of the popula
tion.
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