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~:;;;~1983 ANINUAL REPORT
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BIIG GAME STUDIES
VOLUME V WOLF
Warren B.Ballard
Jackson S.Whitman
Lawrence D.Aumiller
Pauline H-essing
TK
1425
.58
854
00.2324
ALASKA DEPARTMENT OF FISH AND GAME
Submitted to the Alaska Power Authority
April 1984
DOCUMENT No.23'24
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SUSITNA HYDROELECTRIC PROJECT
1983 ANNUAL REPORT
BIG GAME STUDIES
VOLUME V -WOLF
Warren B.Ballard
Jackson S.Whitman
Larry D.Aumi ller
Pauline Hessing
Alaska Department of Fish and Game
Submi tted to the
Alaska Power Authority
April,1984
DEC 3 j ]984
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Y35N
no.131'1
Alaska Resources
Library &Information Services
Anchorage,Alaska
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ANY QUESTIONS OR COMMENTS CONCERNING
'!'IUS RBPOR~SBOtJLD BE D:nu«:T.BD TO
'!JIB ALASltA POWER AtrrBORITY
SUSIDA PROJ~OFPICE
ARLIS
Alaska Resources .
Library &Infonnation Services
Anchorage,Alaska
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PREFACE
In early 1980,the Alaska Department of Fish and Game contracted
with the Alaska Power Authority to collect information useful in
assessing .the impacts of the proposed Susitna Hydroelectric
Project on moose,caribou,wolf,wolverine,black bear,brown
bear and Dall sheep.
The studies were broken into phases which conformed to the antic-
ipated licensing schedule.Phase I studies,January I,1980 to
June 30,1982,were intended to provide information needed to
support a FERC license application.This included general
studies of wildlife populations to determine how each species
used the area and identify potential impact mechani sms.Phase I I
studies began in order to provide additional information during
the anticipated 2 to 3 year period between application and final
FERC approval of the license.Belukha whales were added to the
species being studied.In these annual or final reports,we are
narrowing the focus of our studies to evaluate specific impact
mechanisms,quantify impacts and evaluate mi tigation measures.
This is the second annual report of ongoing Phase II studies.In
some cases,objectives of Phase I were continued to provide a
more complete data base.Therefore,this report is not intended
as a complete assessment of the impacts of the Susi tna Hydro-
electric Project on the selected wildlife species.
The information and conclusions contained in these reports are
incomplete and preliminary in nature and subject to change with
further study.Therefore,information contained in these reports
is not to be quoted or used in any publication without the
wri tten permission of the authors.
The reports are organized into the following 9 volu;nes:
Volume I.
Volume I I.
Volume III.
Volume IV.
Volume V.
Volume VI.
Volume VII.
Volume VIII.
Volume IX.
Big Game Summary Report
Moo se -Downstream
Moose -Upstream
Caribou
Wolf
Black .Bear and Brown Bear
Wolverine
Dall Sheep
Belukha Whale
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SUMMARY
During 1982 and 1983 wolf studies were continued in the Susitna
River Basin to investigate potential impacts of the proposed
Susitna Hydroelectric Project.Between 1 November 1981 and 31
December 1983,42 wolves were captured and outfitted with
transmi tter-equipped collars to enable researchers to document
movements,denning and rendezvous locations,habitat use,and
food habits.Throughout the period,a total of 13 different
packs and 1 lone wolf were known to be using areas in or adjacent
to the Devil Canyon or Watana impoundment zones.During any
particular year 5-6 individual wolf packs used areas to be
inundated by the proposed project.Six hundred forty-nine radio
locations yielded 945 wolf sightings upon which this report is
based.Individual pack histories are presented.
Terri tory sizes of 7 intensively monitored packs ranged from
127 mi 2 to 602 mi 2 (329 km:/.to 1559 km 2 L and averaged 452 mi 2
(1171 km 2 )in 1982 and 1983.Territory sizes varied considerably
among packs,probably due to pack size,prey densities,frequency
of moni toring,and adjacent pack boundaries.
Distribution of wolf packs is suspected to be virtually complete
in the basin,with elevational use generally restricted to less
than 4,000 ft.Elevational distribution varies seasonally and is
probably dependent on relative densities of maj or prey.Both
moose and wolves used lowest annual elevations in February,with
a general increase in elevational use until October with sub-
sequent declines thereafter.
Analyses of food habits of wolves were based largely on aerial
observations of wolves at kills.Moose of all age classes
represented 61%of,the diet,with caribou comprising 30%.
Analysis of 1982 scat collections supported conclusions drawn
from aerial observations.
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Probably the most important impact on wolves resulting from the
proposed project will be lowering of wintering densities of
primary prey species (moose and caribou)in the impoundment zone,
wi th resultant declines in wolf numbers.Secondly,loss of
habitat through inundation and facilities development will
undoubtedly force wolves to readjust territory boundaries
resul ting in intra-specific strife.This will affect not only
wolf packs presently in the basin (especially the Watana pack),
but also packs far removed from the area.
iv
TABLE OF CONTENTS
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SUMMARY
LIST OF TABLES
LIST OF FIGURES
INTRODUCTION
iii
vii
viii
1
METHODS . . . . . . . . . . . . . . . . . . . . . ...2
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RESULTS AND DISCUSSION .
Capture and Telemetry Data .
Individual Pack Descriptions
Broad Pass Pack
B-S Pack . . .
Canyon Creek Wolf 406
Clearwater Pack .
Goose Creek Pack
Jay Creek Pack
Maclaren Wolf 369
Portage CreeR Pack
Snodgrass Lake Pack .
Susitna-Stephan Pack
Susitna II Pack.
Talkeetna River I Pack
Talkeetna River II (Wolf 413)
Watana Pack
Other Packs
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3
3
6
6
6
9
10
10
10
11
12
13
14
14
15
15
15
17
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TABLE OF CONTENTS (cont'd)
Distribution .
Numbers of Wolves and Territories
Elevational Use
Food Habits
Impacts
PROPOSED STUDIES
ACKNOWLEDGEMENTS
LITERATURE CITED
APPENDIX I
vi
Page
18
21
21
25
28
32
33
34
36
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LIST OF TABLES
Table 1.Wolves captured in the upper
Susitna River Basin,Alaska
between November 1981 through
December 1983 . . .
Table 2.Telemetry data and status of
wolves captured in the upper
Susitna River Basin,Alaska
December 1981 through
December 1983
Table 3.Seasonal numbers of wolves per
pack occupying the Susitna River
Basin of southcentral Alaska from
fall 1982 through fall 1983 .
Table 4.Radio locations and territory
sizes of selected wolf packs
in the Susitna River Basin
during 1982 and 1983
Table 5.Kills at which 7 wolf packs were
observed during 1982 and 1983
in the Susitna River Basin of
southcentral Alaska .
vii
Page
4
7
22
24
27
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Fig.1.
Fig.2.
Fig.3.
Fig.4.
Fig.5.
LIST OF FIGURES
Known and suspected pack
boundaries for wolf packs
occupying the Susitna River
Basin from December 1981
through December 1982 . . . . . .
Known and suspected pack
boundaries of wolf packs
occupying the Susitna
River Basin during 1983
Combined monthly numbers of
wolves in 8-9 packs in the
upper Susitna River Basin,1982-83
Watana Pack monthly elevations
and monthly elevation averages
of 9 radio-collared moose within
the Watana Pack territory during
1981 and 1982 in the Susitna River
Basin,Alaska
Use and availability of various
elevational strata by the Watana
Wolf Pack from November 1981
through December 1982 .
viii
Page
19
20
23
26
31
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INTRODUCTION
The proposed Susitna Hydroelectric Project,consisting of 2 dams
and power generating facili ties,is expected to impact a vast
area in the upper Susitna River Basin of southcentral Alaska.A
number of wildlife species occurring within the inundation zones
and along adjacent areas will probably be affected detrimentally.
Of particular importance are the effects on moose (Alces alces)
and caribou (Rangifer tarandus)populations.Subsequently,large
carnivore and scavenging species will also be affected.The
impacts on these carnivorous species which depend on large
ungulates for food could be substantial.Wolves (Canis lupus),
black bears (Ursus americanus),brown bears (Ursus arctos),and
wolverine (Culo gulo)have been identified as the large carni v-
orous species most likely to be severely impacted (Ballard et al.
1982).
Wolves in the upper Susitna River Basin and adjacent areas have
been the focus of study for over 30 years (Ballard et al.1981).
History of Game Management Unit (GMU)13 wolves has been de-
scribed in detail by Ballard et al.(In Review).Wolf studies of
particular importance to the proposed Susitna Project were
reported by Ballard et al.(1982,1983)and were designed to
gather preliminary data on pack sizes,territory boundaries,den
and rendezvous site use,turnover rates,and feeding habits.
This report updates the Phase I report (Ballard et m.1982)and
encompasses the period from October 1981 through December 1983.
Because the information contained in this report treats only
portions of continuing studies,it should not be used in sci-
entific technical publications without the wri tten approval of
the investigators.
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METHODS
Methods used in this study were discussed by Ballard et al.
(1982 i In Review)and will only be briefly reviewed here.
Wolves were captured according to methods described by Ballard et
al.(1982).When practical,blood and hair samples were col-
lected from captured wolves,and the following body measurements
were recorded:weight,body length,tail length,heart girth,
neck circumference,chest height,shoulder height,and length of
canines.
Approximate ages of captured wolves were determined on the basis
of tooth eruption and wear.Trappers and hunters were encouraged
to provide wolf carcasses by offering them $10.00 per carcass,
and an additional $25.00 for turning in radio-collars.Ages of
most harvested wolves were determined by a combination of tooth
eruption and wear,tooth sectioning and counts of cementum annuli
(Goodwin and Ballard 1985)and by examination of epiphyseal
cartilage of the longbone (Rausch 1967).
Captured wolves were fitted with radio-equipped collars manu-
factured by Telonics,Inc.(Mesa,AZ.),and were located period-
ically from fixed-wing aircraft.Radio-collared wolves were
visually observed,when possible,using methods described by Mech
(1974).On each tracking flight the following data were re-
corded:exact location (plotted on 1:63,360 USGS maps),eleva-
tion,aspect,slope,associated wolves (color,ages),activity
(bedded,feeding,etc.),kill data (species,sex,age,percent
consumed,approximate length of time the animal had been dead),
associated species,date and time of observation,and general
habi tat classification according to Viereck and Dyrness (1980)
and Viereck et al.(1982).
The size of pack territories was outlined by plotting all radio
locations on maps,and connecting outermost observations (Mohr
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1947).Radio locations of pack members in the process of di s-
persing were not included.A compensating polar planimeter was
used to determine terri tory size.
Information on wolf food habits was collected from observations
of radio-collared wolves at kill sites.This method grossly
underestimates the precentage of the diet made up of small prey
such as rodents and birds,as the smaller items are consumed
quickly,and are often impossible to identify from aerial obser-
vations.When practical,ungulate kills were examined on the
ground according to methods described by Stephenson and Johnson
(1973)and Ballard et 01.(1979).In most cases,the mandibles
and a femur or metatarsal were collected to aid in establishing
the animal's age (Sergeant and Pimlott 1959,Skoog 1968)and
physical condi tion (Neiland 1970).
Wolf dens and rendezvous sites located through radio-tracking
efforts were inspected in August or September after the wolves
had vacated the sites.All scats were collected,and records of
other food remains were made.Scats were collected and individu-
ally placed in paper bags,autoclaved,and stored for later
analysis .
RESULTS AND DISCUSSION
CAPTURE AND TELEMETRY DATA
Between 1 October 1981 and 31 December 1983,42 wolves were
captured and instrumented in the upper Susitna River Basin
(Table 1).A total of 13 wolf p~cks and 1 lone wolf were known
to be using areas bordering the Devil Canyon or Watana impound-
ment zones during thi s reporting period,although not concur-
rently.Use of the impoundment zones by these packs is dynamic,
as some packs no longer exist (due to harvest and dispersal),and
other packs immigrated into the area and were not located until
3
Table!.(cont'd)
-ACCESSION DATE OF BLOOD PARAMETERS
PACK NAME NUMBER CAPTURE AGE COLOR SEX P.C.V.]j %Hb !)
'~4/10/81IWatana122308 4 gray F 47.0 17.5
122346 4/11/82 3 gray F 30.0 12.3
122400 4/09/82 5+gray M 46.0 17.0,-122401 4/09/82 2 gray M NR NR
122402 4/09/82 pup gray F 49.0 17.5
~1/Packed cell volume
2/%Hemoglobin
3/Not recorded
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ei ther late in 1982 or 1983.During any particular year 5-6
individual wolf packs had territories which overlapped the areas
proposed for inundation.
A total of 649 radio locations were obtained on 42 radio-collared
wolves over a 25 month period (Table 2).These relocations
resul ted in 945 wolf sightings.Descriptions of the dynamics of
each pack are described in the following sections.Histories of
particular packs prior to 1982 were described by Ballard et al.
(1981 and 1982).
INDIVIDUAL PACK DESCRIPTIONS
Broad Pass Pack
An attempt was made in December 1982 to capture and instrument
additional members of the Portage Creek Pack since contact with
most of the pack had been lost in late fall.The Portage Creek
pack often moved to the.East Fork of the Chulitna River and Broad
Pass areas,so an attempt was made to locate them there.On 2
December 1982,a wolf pack containing at least 12 members was
located within the area traditionally used by the Portage Creek
Pack.Four of the pack members were captured and instrumented
(Table 1).Subsequent locations revealed that it was not the
Portage Creek pack but a pack which occupied the area to the
northwest extending into Denali National Park.Because this pack
does not utilize areas to be severely impacted by the proposed
project they were only located on 14 occasions during the 2 year
period (Table 2.)Based upon the limited number of locations the
pack occupied an area of 754 km 2 •
B-S Pack
Throughout 1982,the B-S Pack was comprised of 3 adult members
(numbers 367,368 and 405).They were successful in rai sing a
litter of 4 pups.Because of their differential degree of
6
Table 2.Telemetry data and status of wolves captured in the upper Susitna River
Basin,Alaska from December 1981 through December 1983.
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PACK NAME
ACCESSION NO.RADIO
NUMBER LOCATIONS
NO.WOLF NO.PACK
SIGHTINGS DAYS STATUS (12/31/83)
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Broad Pass
B-S
Canyon Creek
Clearwater
Goose Creek
Jay Creek
Maclaren
Portage Creek
414
415
416
417
367
368
405
479
480
406
323
424
324
423
199
355
356
357
358
365
369
361
362
363
364
403
404
2
14
13
7
52
40
42
9
10
8
4
4
5
8
5
7
7
7
7
5
27
12
5
16
2
18
22
143
188
8
8
15
68
26
100
16
55
8
4
10
8
27
33
Trapped 12/82
Still have contact
Still have contact
Missing 3/5/83
Ground shot 1/84
Killed by Watana wolves 3/83
Ground shot 1/84
Ground shot 12/83
Ground shot 1/84
Ground shot 2/83
Illegal harvest 1/83
Illegal harvest 1/83
Trapped 1/83
Ground shot 2/83
Illegal harvest
Illegal harvest
Illegal harvest
Illegal harv~st
Illegal harvest
Illegal harvest
Killed by other wolves
Snared upper Gakona River 3/83
Suspected illegal harvest
Lost contact 8/18/82
Lost contact 4/10/82
Ground shot 11/83
Ground shot 9/83
Snodgrass Lake 330
336
407
Susitna-Stephan 331
349
30
22
8
9
4
104
9
32
9
Suspected illegal harvest 2/83
Missing
Ground shot 2/83
Shot by ADF&f-Unit 20 control
Suspected illegal harvest
Talkeetna R.I 359
360
-Susitna II 491 3
4
4
20
7
7
3
5
Still have contact
Suspected illegal harvest
unknown
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attentiveness to the den site,it was suspected that wolf 367 was
the producing female,368 was the alpha male,and 405 was a
subordinate member of the pack.Wolves 367 and 368 were instru-
mented in December 1981,and wolf 405 was instrumented in April
1982.At the end of 1982,the pack still contained 7 members.
Due to unknown reasons the adult members of the B-S pack became
separated from the pups in early February.Two pups were cap-
tured and radiocollared in April 1983.The two groups remained
separated until early May when they were observed close to the
1983 den site.
During 1983 the pack moved further west apparently in response to
the vacancy created by the elimination of the Jay Creek pack.An
eastward expansion of the Watana wolves also occurred at this
time,resulting in a confrontation with the B-S pack which
resulted in the death of B.S.wolf 368.
The pack was located on only 3 occasions between 27 May and 31
December 1983.By early December the pack numbered 8 and had
obviously produced pups.Either some wolves had dispersed or the
1983 litter size was small because based on average GMU-13 litter
sizes the pack should have numbered 12.From December 1981
through December 1983 the pack occupied on area of 1412 km 2 •In
late December and early January 1984 the entire pack was killed
by aircraft-assisted hunters.
Canyon Creek Wolf #406
In October 1982,a lone yearling gray female (406)was captured
and instrumented in the upper Susitna River Basin on Canyon Creek
near the Denali Highway.This single wolf was locat-.ed on 8
occasions between October 1982 and late February 1983 (Table 2)
and ranged from Deadman Creek to the east fork of the Jack River
to Canyon Creek.This suggests that during this time period the
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area immediately north of the Watana Pack was probably vacant
except for sporatic excursions by the Snodgrass Lake pack and
this single wolf.Based upon the limited number of relocations
the single wolf occupied an area of 671 km 2.
Clearwater Pack
Contact with the Clearwater Pack was established in December 1982
when Watana wolf 323 was relocated between Clearwater Creek and
the Maclaren River.Wolf 323 was accompanied by another wolf
which was captured and instrumented on 12 December 1982 (adult
gray male 424).Only 4 locations were gathered on this pair
before they were killed by aircraft-assisted hunters.This pair
was apparently recolonizing the area left vacant after the death
of wolf 369 (see Maclaren wolf history).Because of the small
number of locations no terri tory size was computed.
Goose Creek Pack
Contact with the Goose Creek Pack was established on December
1982 when Watana wolf 324 (adult gray female)was recaptured and
her radio-collar replaced.Her lone associate was captured
12 December 1982 (adult gray male #423).The wolves appeared to
be in the process of colonizing the area but were killed in early
1983.Because of the small number of relocations the estimated
terri tory size of 322 kIn 2 was probably low.
Jay Creek Pack
During winter 1979-80 there were reports of 10 to 11 wolves
wi thin the Jay Creek pack area.Six wolves were ground shot at
that time.At least 4 wolves were observed on the Susitna River
near Jay Creek during a moose survey in March 1980,indicating
that the pack area was still occupied,probably by descendants of
wolf 199.No further information on this pack was obtained until
November 1981.
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Radio contact with the Jay Creek Pack was first established on 30
November 1981.Prior to contact,track sightings and uncollared
wolf sightings in the area suggested the pack's presence for
quite some time.On 30 November,the pack contained 12 members
and wolves 355, 356,357 and 358 were captured and instrumented
(all pups;male,female,female and male,respectively).On 6
December,2 additional wolves were captured (adult gray females
199 and 365).Wolf 199,originally from the Tsusena Pack,was
the alpha female.The Jay Creek Pack inhabited the area from
Coal Creek to the Susitna River and Jay Creek without attrition
until mid-January when they were suspected to have been illegally
shot by aerial hunters.Four wolf carcasses were found on the
Susitna River in early March,and it was suspected that the rest
had also been killed.The unskinned carcass of wolf 365 was
found in July and examination of the carcass revealed punctures
in the pelvis which matched the size of OO-buck shotgun pellets.
Maclaren Wolf 369
In early December 1981 an adult black female (#369)was captured
and radio-collared near Clearwater Creek.At capture,she was
accompanied by 1 gray wolf which was apparently harvested in
early January.Regular radio-contact with 369 was continued
until 30 October 1982,when she was suspected to be sick or
injured.Her carcass was retrieved in early 1983 and indications
were that she had been attacked by other wolves,was weakened,
and later starved to death.Her territory size during the period
she was instrumented was 127 mi 2 (329 km 2 ) •
It was interesting to note that during the time she was being
monitored,wolf 369 showed fidelity to a den site,although no
other wolves were seen at the den and no pups were kn?wn to have
been produced.
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Portage Creek Pack
The Portage Creek Pack was first located on 2 December 1981,and
4 members were captured and radio-collared (wolf 361 -adult gray
male;362 -adult gray female;363 and 364 -pup gray females).
At that time,there were only 5 individuals in the pack.Reloca-
tion of the pack was not successful on 9 succeeding attempts,and
it was not until 14 January 1982 that they were found.By then,
the pack contained at least 8 members.
By early March the pack had split up,with a maximum of 5 wolves
observed together.Wolf 362 was suspected to have been illegally
shot from the air,as the collar was retrieved and had obviously
been cut off and the harvest was never officially documented on
mandatory sealing certificates.Two additional wolves were
captured and instrumented in April 1982 (403 -adult gray female;
404 pup gray female)bringing the Portage Creek Pack to 5
members,all of which were instrumented.
Wolf 364 was suspected to have dispersed or was harvested from
the pack in mid-April,because she was never again observed with
the pack.The den site was located on 14 June at which time the
pack was comprised of 2 adults and 2 yearlings (adults 361 and
403;yearlings 363 and 404).At collaring wolf 403 was pregnant,
thus was assumed to be the alpha female.The den site was
inspected on 17 September and several pup scats were collected
even though no pups were ever observed.
By late fall 1982,radio contact with all but 2 members of the
pack was lost.We suspected that several of the wolves had been
shot or dispersed.Radio-collared wolves 403 and 404 and 2
uncoll~red wolves were the only pack members known to be still
alive.On March 1983,wolf 361 was snared along with 1 associate
on the upper Gakona River,a straight1ine movement of 180 km.
The pack denned at an unknown location in 1983.They were not
12
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Pups began
area from....the fall.
mi 2 (1559
located between early May and December 1983.Wolf 403 was killed
in November while wolf 404 was shot in September 1983.In early
fall 1983,the pack had numbered 8 according to public reports.
Snodgrass Lake Pack
Wolf 330 apparently colonized the Snodgrass Lake Pack in November
1981 after dispersing from the Susitna Pack in early May.This
2-year-old gray male was accompanied by another wolf which was
captured and instrumented on 6 December 1981 (2-year-old gray-
black female wolf 366).They remained together through the
winter and were observed at a den site near Butte Creek where at
least 6 pups were produced .
to travel with the adults by mid-September and used an
upper Brushkana Creek to Clearwater Creek throughout
Total area encompassed within their territory was 602
km 2).On 15 October 1982,the 7-member pack·was
located and an additional wolf (pup gray-brown male wolf 407)was
instrumented.
On 18 or 19 November,a trapper near Susi tna Lodge watched an
airplane harassing the pack,and subsequently observed the
occupant(s)aerially shooting at the pack.He reported that they
illegally harvested at least three of the pack members.On
subsequent radio-tracking flights,the pack was found to contain
only 4 members,including wolves 330 and 407.It was not certain
whether wolf 366 had been harvested or her transmitter had
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fai led.In early February 1983,
killed by illegal aerial hunters.
be vacant through~ut 1983.
13
the remainder of the pack was
The pack area was believed to
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Susi tna -Stephan Pack
During the short time the pack existed it was comprised of 2
collared wolves.Wolf 331,originally from the Susitna Pack,had
a hi story of wanderings over much of the upper Susi tna Basin
(Ballard et al.1982).They had apparently not established a
territory during the time they were monitored,as their movements
were typical of dispersing wolves (op.cit.).
Wolf 331 was recaptured along with his lone associate (wolf 349,
adul t gray male)in November 1981 near Stephan Lake.It was
suspected that wolf 349 was either another Susitna Pack member or
was from the Watana Pack (Ballard et al.1982,page 19).
-
On 13 December 1981,wolf 349
legally shot from the air,as
and blood in the snow.This
disperse out of the area.
was suspected to have been il-
evidenced by airplane ski tracks
may have prompted wolf 331 to
On 16 April 1982,ADF&G personnel from the Fairbanks office
reported the death of wolf 331.He had been harvested by
Department personnel as part of control efforts in Game Manage-
ment Unit 20 near Healy;a straightline movement of 129 km.When
killed he was accompanied by 6 wolves,apparently having been
accepted into an exi sting pack.
Susi tna I I
During late June 1983,three wolves were observed near the old
Susitna pack rendezvous site.Since November 1981 this pack area
had been largely vacant except for colonizing pairs.One wolf
(adult female 491)was captured and radio-collared.The pack
produced 6 pups,however no information was obtained on pack
distribution due to inadequate funding.
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Talkeetna River I Pack
Contact with this pack was established in early December 1981.
The origin of the wolves was unknown but because only 2 wolves
(359,adult gray male and 360,adult black female)were present,
they probably had recently colonized the area.They were located
only 3 times between collaring and 4 January 1982,and were
always together.On 16 February,however,only wolf 360 was
seen,,and the signal from 359 was located in steep,rocky terrain
not typical of wolves at this time of year.At this time,wolf
360 appeared to be shy of the airplane,leading us to assume that
they had been shot at by hunters.Because the signal from 359
remainded at that same location on subsequent flights,it was
assumed that aerial hunters had shot him,and the transmitter had
been thrown out of the plane.Wolf 360 was never again located,
and it was assumed that she had either been harvested or had
dispersed.No estimate of territory size was calculated because
of the minimal number of relocations .
Talkeetna River I I
Wolf 413 (adult gray male)was captured on 3 December 1982 on the
Talkeetna River upstream of Prairie Creek.Tracks in the area
indicated other wolves were present.Wolf 413 was observed with
1 other black wolf in early March 1983 and by early April was
associated with 2 blacks.An adult pregnant female was killed as
a result of capture activities in early April.An additional
black adult male (477)was captured and radio-collared at that
time.The pair of males occupied an area of 2081 krn 2 during
1983.
Watana Pack
At the beginning of 1982,the Watana Pack was comprised of at
least 10 members,at least 6 of which were adults.The Watana
Pack resided in a location which may have been beneficial to
15
,,,,,,,",
I
-
-
....
....
their survival.The area has few lakes and much of the area is
heavily timbered so hunter accessibility is limited.In addi-
tion,the"constant human habitation of Susitna Hydroelectric
Project's Watana Camp from 1980-82 and associated aircraft
frequenting the area may have deterred illegal hunters from using
the area.Whether these factors or others were responsible,the
Watana Pack has hi storically realized little attrition and has
been responsible for production of many wolves which have dis-
persed to other areas of southcentral Alaska.
Only 1 Watana wolf was suspected to have been harvested during
1982.This was a wolf found 22 March on the Susitna River,
suspected to have been illegally shot from aircraft.
During 1982 at least 4 radio-collared wolves were known to
disperse from the Watana Pack,either joining existing packs or
starting new packs.We suspect the reason for the high incidence
of dispersal was due to the low attrition wi thin the pack,
resul ting in high pack numbers and subsequent di spersals.In
previous years,at least 5 other wolves from the Watana Pack were
known to have di spersed to new areas.
During recollaring operations in early April 1982,wolf 309 was
accidentally killed.When a necropsy was performed,it was
learned that this wolf was pregnant.Beginning in early May,
however,the Watana wolves were repeatedly located at the den
site of 1981,indicating that a litter had been produced.This
was later verified.Had wolf 309 not been killed,it is sus-
pected that the pack would have double-denned,producing 2
litters .
Pups were first observed in early July 1982.By the end of
summer,the Watana Pack contained at least 18 members,when the
pack was beginning to split up.By late October,there were at
least three groups,two of which were thought to be dispersing .
Wolves 308,400,401 and 402,along with the pups,remained in
16
.....
-
.....
~
I
~
I
the Watana territory.Wolf 346 and two associates moved approxi-
mately 30 mi (48 km)to the upper Coal Creek-Big Bend of the
Susi tna area and was shot in late February 1983.Wolf 323 and
one associate had moved about 43 mi (69 km)to the Round Mountain
area,between the Clearwater and Maclaren Rivers and were prob-
ab1y'attempting to colonize the area after the death of Maclaren
wolf 369.In December,the wolves remained separated and in
thei r respective areas.
Additionally,wolf 324 was captured during the December collaring
operation in the vicinity of Goose Creek-Oshetna River.This
wolf dispersed from the Watana Pack in late April 1982.Its
distance from the Watana den site to recapture location was
approximately 26 miles (42 km).Also sometime between October
1982 and January 1983 wolf 402 dispersed from the Watana area and
had joined a pack of 4 wolves which occupied an area south of the
Alphabet Hills.This latter wolf pack is referred to as the Dog
Lake Pack.By mid-March 1983 the Watana pack had been reduced to
7 wolves due to dispersal and hunting mortality.The pack had an
encounter with the B-S pack on the Susitna River and 1 B-S wolf
was killed.Although the pack denned in 1983 at the same site it
(
had used since 1980,no estimate of the number of pups born was
obtained until early December 1983.At that time the pack
numbered 15.During 1982 and 1983 the pack occupied an area of
1913km 2 •
Other Packs
In addition to the above-mentioned packs,other packs not
directly associated with the impoundment zones were sporadically
moni tored during 1982.These packs were usually located by
following dispersing wolves out of an existi.,g Susi tna Basin
pack,and subsequently either initiating new packs or joining
exi sting packs.
17
,....
As mentioned in the individual pack descriptions,many members of
the Watana Pack left that terri tory and dispersed into vacant
areas or di spersed from the Watana terri tory to join exi sting
packs.In addition to the Clearwater and Goose Creek Packs that
were colonized during 1982,2 other packs have resulted from
dispersals out of the Watana Pack.
The Eureka Pack,inhabiting an area from upper Tyone River to
Eureka to the south of the Susi tna Basin,was colonized by a
Watana wolf (344)prior to 1982.This wolf was accompanied by 2
suspected female wolves in 1982,and they were successful in an
apparent double-denning attempt and raised a total of 13 pups.
It was suspected that the entire pack was killed by hunters and
trappers during 1982-83.
Wolf 325,another Watana wolf,dispersed to the southeast in
1980,and became integrated into the Nelchina Pack.Accompanied
by at least 3 addi tional adults,this pack was successful in
raising 5 pups in 1982.Wolf 325 and several other pack members
were trapped or shot during 1982-83.By late spring 1983,the
pack was comprised of 6 wolves but they did not den and no pups
were produced.
During summer 1983 archeologists reported observing a single wolf
at a den si te near Kosina Creek.Apparently,a pair of wolves
colonized the area south of Jay Creek and successfully produced
pups.Five wolves were observed during late fall 1983.We
suspected that most of the wolves from this pack were killed
during the winter of 1983-84.
DISTRIBUTION
Figures 1 and 2 depict the distribution of known and suspected
wolf packs in the middle Susi tna River Basin during 1982 and
1983,respectively.A minimum of 6 wolf packs utilize portions
of the area to be inundated by the 2 impoundments.The figures
18
Depictions of wolf pack boundaries were deleted from this report
due to previous instances of heavy harvests by individuals using
-
the maps to concentrate their hunting efforts.
available to Susitna personnel upon request.
The data is
r-
I Figure 1.Known and suspected pack boundaries of wolf packs
occupying the Susitna River Basin from December 1981
through December 1982.
19
.....
-
.-
-
-
Depictions of wolf pack boundaries were deleted from this report
due to previous instances of heavy harvests by individuals using
the maps to concentrate their hunting efforts.The data is
available to Susitna personnel upon request .
Figure 2.Known and suspected pack boundaries of wolf packs
occupying the Susitna River Basin during 1983.
20
-
.....
-
.....
also depict the colonization and establishment of new packs
following elimination of some packs due to hunting and trapping.
NUMBERS OF WOLVES AND TERRITORIES
The number of wolves inhabiting areas which could be impacted by
the proposed project has fluctuated from 25 in spring 1983 to 47
in fall 1983 (Table 3).Both hunting and trapping have regulated
the number of wolves occupying the area.Mostly wolf mortality
occurred during the months of January through April primarily
from aircraft assisted ground shooting (Table 2,Fig.3).
Territory sizes of 9 wolf packs in the Susitna River Basin ranged
from 124 mi 2 to 803 mi 2 (322 km 2 to 2081 km 2 ),and averaged 452
mi 2 (1171 km 2 )(Table 4).Some territory sizes may not be
adequately described because some packs have only been located a
few times.
Variation in observed teritory size for individual packs was due
to adjacent pack boundaries,changes in distribution of prey,
number &frequency of monitoring,and increases or decreases in
pack size as a result of hunting mortality,dispersal,and
natality.When a territory is vacant of wolves due to harvest or--natural mortality,it is likely that that area will be recolon-
ized by dispersing wolves within a short period of time (in most
cases,less than a year)or will be usurped by an existing pack.
ELEVATIONAL USE
Generally,wolves restrict their movements to elevations less
than 4,000 ft/1300 m.For example,the Watana Pack had only 2 of
56 (3.6%)observations at elevations great~r than 4,000 ft/1300 m
elevation in 1982.
21
r
-
,~
-
Table 3.Seasonal numbers of wolves per pack occupying the Susitna River Basin of
southcentra1 Alaska from fall 1982 through fall 1983.
No.of wolves
Pack Name Fall 1982 Spring 1983 Fall 1983
B-S 7 6 8
Canyon Creek 1 0 0
Clearwater 2 0 0
East Fork 3 3 3
Goose Creek II 2 0 0
Jay Creek -Clarence Lake 0 >2 5
Maclaren 1 0 0
Portage Creek 4 3 8
Snodgrass 7 0 0
Susitna II ?3 9
Talkeetna II 3 2 2
Watana 16 6 12
Total 46 >25 47
22
I""'(/J
a:40
W
a:l
::E
'~::»3 0
Z
LI.
..J
0 20
==--
I"""
I
-
.....MONT-HlYEAR
-
.-
Figure 3.Combined monthly numbera of wolves In 8-8 packa In the SusJtna
River Baaln.t882-83.
23
f"'"
Table 4.Numbers of radio locations and territory sizes of selected wolf
packs in the Susitna River Basin during 1982 and 1983.
No.No.No.
Radioed of Radio Territory Pack
Pack Name Wolves Locations (mi 2 )(km 2 )Days
Broad Pass 4 36 291 754 16
B-S 5 153 545 1412 55
Canyon Creek 1 8 259 671 8
,....Clearwater 2 8 4
Goose Creek 2 13 124 322 10....
Jay Creek 6 38 8
Maclaren 1 27 127 329 27
Portage Creek 6 75 580 1501 33
....Snodgrass Lake 3 60 602 1559 32
Susitna-Stephan 2 13 9
Talkeetna I 2 8 5
Talkeetna II 3 17 803 2081 13
'Watana 10 190 738 1913 49
Susitna II 1 3 3
48 649 x =452 1171 272
.-
.....
-24
-.
-
.....
Elevational distribution of"wolves varies seasonally.We sus-
pected that the changes in distribution were largely dependent on
changes in prey distribution.For example,the Watana wolves
rely heavily on moose (Ballard et al.1982)as a source of prey
and it was suspected that their differential elevational use was
a reflection of availability of those moose.Mean monthly
elevations of nine radio-collared moose that had annual home
ranges that overlapped portions of the Watana pack territory were
compared with elevations of the Watana Pack (Fig.4).Lowest
mean elevation for both wolves and moose was in February,with
both species exhibiting a general increase in elevations occupied
until October,with subsequent declines thereafter.This suggests
that wolves were following the movements of their principal prey.
FOOD HABITS
Seven radio-collared wolf packs were observed on 76 kills during
1982 and 1983 (Table 5).Moose of all age classes accounted for
61%of the observed prey.Of the moose kills which could be
assigned to 1 of 3 age classes (calf,yearling or adult)both
adult and calf moose comprised 44 and 41%of the kills,respec-
tively.Calf moose do not appear to be preferred prey in early
summer but from November through April they represent a dispro-
portionate portion of the diet (Ballardet al.1981,1982,In
Review).
Caribou comprised the second most important wolf prey species,
accounting for 30%of the observed kills.There appeared to be
no selection for calf caribou.
Aerial observations of wolf food habits during summer were
supplemented with data obtained from scat analyses.Scats were
collected from 5 home sites (includes den and rendezvous sites)
in 1982 (Appendix I).No scats were collected in 1983 .
25
3000
--moose
-wolves
......
/~\
/\
//\
I \
I ~
I
I
.....1/--
/
/
/
I
I
I
I
I
,I
,I,/
,I
'__J
2200
2800
2300
2400
2900
2100
2500
2800
2700
t-
e(
>
W
.J
W
t-
W
W
LL.
z
o
"'"
,~
,~
I"'"
2000 ---...-...-1----...+--+0--+--......-....-+-0-1----+-...--+
M 0 NTH
FIgure -4.Watana Pack monthly elevations and monthly elevation averages
of nine radIo-collared moose within the Watana Pack terrl.tory during 1981
and 1982 In the Sualtna River Basin,Alaska.
26
}I J J )1 I )1 1 ,J I J
Table 5.Kills at which 7 wolf packs were observed during 1982 and 1983 in the Susitna River Basin
of southcentral Alaska.
Pack Name
B-S Fish Lk.Jay Cr.Portage Snodgrass Stephan Watana Total
No.%No.%No.%No.%No.%No.%No.%No.%
Moose.Adult 3 16 2 8 2 100 1 20 1 14 ----3 20 12 16
Moose,Yrlg.1 5 2 8 --------I 14 --------4 5
Moose.Calf ,4 21 2 8 ----I 20 ----I 50 3 20 11 14
Moose,Unknown 5 26 4 15 ----2 40 2 29 ----6 40 19 25
Total Moose 13 68 10 38 2 100 4 80 4 57 1 50 12 80 46 61
Caribou.Adult 2 11 11 42 --------2 29 1 50 3 20 19 25
N.......Caribou.Calf 1 14 1 1------------------------
Caribou.Unknown ----3 12 ------'--------------3 4
Total Caribou 2 11 14 54 --------3 43 1 50 3 20 23 30
Ungulate,sp.unk.3 16 ------------------------3 4
Sheep.Adult ------------I 20 ------------I 1
Total Ungulate 18 95 24 92 '2 100 5 100 7 100 2 100 15 100 73 .96
Other:Beaver ----I 4 --------------------I 1
Snowshoe ----I 4 --------------------I 1
Unknown 1 5 ------------------------I 1
Total Other 1 5 2 8 --------------------3 4
Grand Total 19 100 26 100 2 100 5 100 7 100 2 100 15 100 76 100
-
Incidence of moose and caribou in the wolf diet is probably a
reflection of availability due to population numbers.The
Nelchinacaribou herd reached a low of approximately 10,000
animals in 1973 (Pitcher,pers.comm.).By 1981,the population
had increased to approximately 22,000 (op.cit.).We suspect the
increase in the caribou population has made them generally more
available to wolves.If true,this would suggest that as the
herd grows,caribou will probably become more important in the
diet.This may be beneficial to moose populations in the Basin
if caribou populations become high enough to provide the maj or
source of food for wolves.
IMPACTS
Impacts of the proposed impoundments and their associated borrow
pits,transmission corridors,work camps and facilities are
difficul t to quantify at this time.However,based on earlier
research and that reported herein,some impacts can be estimated.
Probably the most significant impact the impoundments will have
on wolves will result from a change in population density,
distribution,sex and age composition and/or physical condition
of moose and caribou.The majority of the wolf's diet in this
area is moose,and any decrease in prey numbers will probably be
reflected in both wolf density and distribution.Ballard et al.
(1984)estimated that approximately 2,800 moose will be directly
impacted by the impoundments.During the impoundment filling
stage and for at least 1 year following inundation,there will
probably be an increase in wolf numbers in response to higher
concentrations of moose adjacent to the impoundments.There will
be a high number of displaced moose which wi).l be concentrated
adjacent to the reservoirs due to the decreased availability of
usable habitat.However,the duration of this relatively high
moose concentration will probably be short,i.e.2-3 years.
28
....
...,.
No quantitative data are presently available upon which to base
current moose carrying capacity figures.However,our obser-
vations of browse utilization in areas presently used by moose
for winter habitat indicates that preferred willow (Salix spp.)
species are heavily used in many areas.Displacement of moose
from the low elevation areas to available habitat above high pool
level will undoubtedly lead to a further increase in the use of
preferred areas,ultimately leading to a deterioration of that
preferred habitat.Ultimately,remaining wolves and the area J s
abili ty to support larger numbers will be reduced,but for a
short period they will remain relatively high and further depress
the moose population and possibly prevent it from increasing.
For the period of 1972-73 through 1982-83,mandatory wolf sealing
documents have shown that 48 percent of all harvest in GMU-13 is
from trapping and snaring.Land-and-shoot methods accounted for
42 percent of the harvest.An increase in access roads and the
proposed permanent village for project personnel will result in a
significant increase in human use of the area.Correspondingly,
there will probably be a higher incidence of hunting,trapping
and accidental mortality upon wolves.Indirect effects upon
wolves resulting from higher human populations will probably also
occur.Acti vi ty near den and rendezvous sites in early summer
will certainly disrupt,and in some cases,will probably cause
wolves to abandon den and feeding sites.Den site abandonment
could lead to higher pup mortali ty.
Inundation of den and rendezvous sites,travel corridors and
hunting/feeding areas will eliminate portions of wolf terri-
tories.Loss of this habitat will force wolves to readjust
territory boundaries and will probably result in increased inter-
pack strife.Since present wolf mortality from some of the packs
adjacent to the Susitna River are low (Watana Pack in particular)
with subsequent high dispersals to surrounding areas,this area
acts as a reservoir in supplying wolves to adjacent areas .
Should mortality within these packs increase,there will probably
29
.-
-
-
be less dispersal away from the area.Consequently,the reduc-
tion of wolf numbers adjacent to the impoundments may well affect
not only those packs immediately adjacent to the river,but also
packs far removed from the area.Movements of over 50 mi les
(80 km)away from the Watana terri tory by wolves which either
joined new packs or initiated new packs have been recorded.
Of the 6 packs which will be impacted by the proposed project,
the Watana pack will be one of the packs to be impacted most
severely.The effects of habitat inundation on the Watana wolves
was selected for a further analysis because of the relatively
high number of relocations.During 1982 the Watana Pack occupied
a territory of 482 mi 2 (1246 km 2 )within and adjacent to both the
proposed Watana and Devil Canyon impoundments.Twenty-six of 58
(45%)relocations of Watana Pack members were at or below high
pool level of the impoundments (Fig.5).During the first half
of the year (January through June)over half (57%)the recorded
observations were at or below maximum pool level.Fifty-one mi 2
of the 482 mi 2 territory were greater than 4,000 ft.altitude and
were rarely used by the pack «4%of all relocations).Of the
431 mi 2 of usable habitat wi thin the Watana pack terri tory,55
mi 2 would be inundated by the two impoundments.However,the 55
mi 2 (13%of territory)accounted for 45%of the annual locations
of the pack,indicating that the lower elevations are preferred
by wolves in the Susi tna River Basin.This was probably the
resul t of higher concentrations of moose in these lower al ti-
tudinal areas.
Preliminary borrow and encampment site locations have been
delineated and their locations will further limit the extent of
the Watana terri tory.The exact percent of habitat loss of the
Watana territory is not known.In particular,quarry sites A and
B,and borrow sites D,E,F,I,J,and L will at least have
portions within the Watana territory and will impact the Watana
wolves during and/or after construction of Watana Dam.
30
--j 11I J J ])I 1 J }
22
20
18
18
t-14
Z
W 12
0
a:
w w 10
I-'0-
8
8
4
2
0 '1400
1600
ELEVATION
o percent available
t~5Iwl percent used
4~1142.01
4600 4800
51.01
5600
Fillure 6;U~e and avaliablUtv o'varlo ...a elevatlonal atrata bV the Wa'ana Wolf PaCk from November 1881
through December 1982,
.....
r
The percent of various elevational strata available to the pack
(calculated by random selection of 482 points within the Watana
terri tory)compared to the percent of radio-locations at those
various elevations shows that between 1801 and 2200 ft.were
significantly preferred elevations (Chi-Square test,P<O.OOS)
(Fig.5).These elevational strata were available to Watana
wolves in 13 percent of their terri tory,yet were used on 45
percent of the locations.The inundation of this zone will
undoubtedly affect the shape and extent of wolf territories and
subsequent recruitment and mortality of wolves in the Susi tna
Basin.
PROPOSED STUDIES
Continuation of monitoring efforts of wolves already instrumented
is important.Population and individual pack boundaries are
highly dynamic and documentation of denning,rendezvous,and
hunting/traveling areas should be continued.Efforts should be
made to instrument numbers of additional packs,especially in the
area from Devil Canyon to Sherman.Of particular concern is a
better representation of preferred habitat types when higher
resolution vegetation mapping is finished.
In conjunction with ongoing moose and caribou studies,investi-
gations of calf consumption by wolves should be conducted.More
accurate documentation of food habits could be gathered in this
way.
Mitigation of the losses of major prey species (moose and cari-
bou)is of major importance to the continued viability of wolf
populations.Evaluation of those mitigation options include their
impacts on wolves .
32
.....
-
ACKNOWLEDGEMENTS
ADF&G personnel Sterling Miller,Dennis McAllister,Enid Goodwin,
John Westlund,Jim Dau and Craig Gardner participated in one or
more aspects of the pro j ect .
Without the help and continued cooperation of a number of pilots,
this study would not have been possible.Ai and Jerry Lee,Lee's
Air Taxi Service,Harley and Chuck McMahan,Ken Bunch and Don
Doering all contributed during collaring and/or radio-tracking
efforts.Vern and Craig Lofstedt,Kenai Air Service,supplied
their skills during collaring operations by not only piloting the
helicopters,but participating in the processing of drugged
animals.
Special thanks go to Karl Schneider,ADF&G,for his guidance and
support throughout the project and for his editorial comments on
the report.Nancy Tankersley also provided valuable editorial
assistance.Susan Lawler provided typing assistance.
33
--
-
LITERATURE CITED
Ballard,W.B.,A.W.Franzmann,K.P.Taylor,T.Spraker,C.C.
Schwartz and R.O.Peterson.1979.Comparison of tech-
niques utilized to determine moose calf mortality in Alaska.
Proc.N.Am.Moose Coni.Workshop,Kenai,Alaska.(5):362-
387.
Ballard,W.B.,R.o.Stephenson and T.H.Spraker.1981.
china Basin Wolf Studies.Alaska Dept.Fish and Game.
Proj.Final Report,W-17-9 and W-17-10.
Nel-
P-R
Ballard,W.B.,C.L.Gardner,
1982.Susitna Hydroelectric
Volume V -Wolf.220 pp.
J.H.Westlund and J.R.Dau.
Pro j ect Phase I Final Report.
-
Ballard,W.B.,J.S.Whitman,L.D.Aumiller and P.
1983.Susitna Hydroelectric Project Phase II
Report.Vol.III Moose-Upstream.61pp.
Hessing.
Progress
.....
Ballard,W.B.,J.S.Whitman,L.D.Aumiller and P.Hessing.
1984.Susitna Hydroelectric Project Annual Progress Report.
Upstream Moose.
Ballard,W.B.,S.M.Miller,K.B.Schneider,and R.o.
Stephenson.In Review.Ecology and relationships of
wolves,brown bear and moose in southcentral Alaska.Wildl.
Monograph.
Goodwin,E.and W.B.Ballard.1985.Use of tooth cementum for
age determination of gray wolves.J.Wildl.Manage:in
press.
r
Mech,L.D.1974.Current techniques in the
wilderness carnivores.Proc.XI Internat.
Biol.P.315-322.
34
study of elusive
Congress of Game
Mohr,C.O.1947.Table of equivalent populations of North
American small mammals.Am.MidI.Nat.37(1):223-249.
Neiland,K.A.1970.
in caribou femurs.
Weight of dried marrow as indicator of fat
J.Wi ldI.Manage.34 (4):904-907 .
Rausch,R.A.1967.
wolves,Alaska.
Some aspects of the population ecology of
Am.Zool.7:253-265.
Sergeant,D.E.and D.H.Pimlott.
moose from sectioned incisor
23(3):315-321.
1959.
teeth.
Age determination in
J.Wildl.Manage.
Skoog,R.O.1968.
granti)in Alaska.
California.699 pp.
Ecology of
PhD.Thesis,
caribou
Univ.
(Rangifer tarandus
of Cal.,Berkeley,
Stephenson,R.O.and L.
ment of Fish and Game.
Johnson.Wolf Report.
P-RProj.Rept.,W-17-4.
Alaska Depart-
52pp.
Viereck,L.A.and C.T.
fication system for
Service.Pac.N.W.
PNW-106.
Dyrness.1980.A preliminary classi-
vegetation of Alaska.U.S.D.A.Forest
Forest and Range Expt.Sta.Report.
Viereck,L.A.,C.T.Dyrness,and A.R.Batten.1982.1982
Revision of Preliminary Classification for Vegetation of
-Alaska.Proc.of Workshop on Classificaton of Alaskan
vegetation.72 pp.
35
APPENDIX I.WOLF SCAT ANALYSIS RESULTS.
Portage Creek 1982 Densite
PUP
(N =51)
ADULT
(N =38)
TOTAL
(N =89)
Prey Species N %N %N %
-unidentified birds 6 5.9 2 2.0 8 7.9~,:;
vegetation 7 6.9 7 6.9
F"'"
TOTAL 59 42 101
items/scat 1.16 1.11 1.13
~
-36
.....
!
....Appendix I.(cont I d)
~
B.S.Lakes 1982 Densite
PUP ADULT TOTAL
(N =40)(N =12)(N =52 )
Prey Species N %N %N %
~
adult moose 13 23.2 13 23.2
calf moose 17 30.4 4 7.1 21 37.5
total moose 30 53.6 4 7.1 34 60.7
r-caribou 4 7.1 4 7.1 8 14.3!
total ungulates 34 60.7 8 14.3 42 75.0
beaver 2 3.6 2 3.6 4 7.1
microtine rodent 2 3.6 1 1.8 3 5.4
ground squirrel 1 1.8 1 1.8 2 3.6
.-porcupine 1 1.8 1 1.8
total mammals 40 71.4 12 21.4 52 92.9
"'"
.-
""""I
unidentified birds
TOTAL
items/scat
2
42
1.05
3.6
37
2
14
1.17
3.6 4
56
1.08
7.1
r Appendix I.(cont I d)
~
Snodgrass Lake 1982 Densite
PUP ADULT TOTAL
(N =140)(N =17)(N =157)
Prey Species N %N %N %
-adult moose 27 15.6 5 2.9 32 18.5
~calf moose 6 3.5 6 3.5 12 6.9
total moose 33 19.1 11 6.4 44 25.4
caribou 46 26.6 2 1.2 48 27.7
total ungulates 79 45.7 13 7.5 92 53.2
beaver 45 26.0 6 3.5 51 29.5
micro tine rodent 3 1.7 6 3.5 9 5.2
~~
snowshoe hare 8 4.6 8 4.6
ground squirrel 7 4.0 7 4.0
porcupine 5 2.9 5 2.9
total mammals 147 85.0 25 14.5 172 99.4
ptarmigan
TOTAL
items/scat
1
148
1.06
0.6
38
25
1.47
1
173
1.10
0.6
-Appendix I.(cont'd)
r-
Watana 1982 Densite
PUP ADULT TOTAL
(N =94)(N =57)(N =151)-Prey Species N %N %N %
~
adult moose 34 20.9 20 12.3 54 33.2
~calf moose 27 16.6 11 6.7 38 23.3
total moose 61 37.4 31 19.0 92 56.4
caribou 24 14.7 17 10.4 41 25.2
unknown ungulate 1 0.6 1 0.6 2 1.2-total ungulates 86 52'.8 49 30.1 135 82.8
snowshoe hare 4 2.5 4 2.5 8 5.0r-
beaver 3 1.8 2 1.2 5 3.0-microtine rodent 4 2.5 4 2.5
ground squirrel 1 0.6 1 0.6
total mammals 97 59.5 56 34.4 153 93.9
~,
unidentified birds 4 2.5 1 0.6 5 3.1
vegetation 2 1.2 2 1.2 4 2.4
wolf hair (grooming -1 0.6 1 0.6
_..;;;,T...;;.O..;;;,T;;;.;A;;:;;L ~;;..10~3.;....-6~O;...._.....;1;;;..6,;,...3;....__
items/scat 1.10
39
1.05 1.08
r
Appendix I.(cont'd)
Watana 1982 Rendezvous Site
PUP
(N =40)
ADULT
(N =18)
TOTAL
(N =58)
Prey Species N %N %N %
adult moose 13 20.6 6 9.5 19 30.2
calf moose 2 3.2 2 3.2 4 6.3
total moose 15 23.8 8 12.7 23 36.5
~caribou 14 22.2 8 12.7 22 34.9
unidentified ungulate 2 3.2 1 1.6 3 4.8.....
total ungulates 31 49.2 17 27.0 48 76.2
snowshoe hare 2 3.2 1 1.6 3 4.8
marten 2 3.2 2 3.2
red squirrel 1 1.6 1 1.6
microtine rodent 1 1.6 1 1.6
ground squirrel 1 1.6 1 1.6
unknown mammal 1 1.6 1 1.6
total mammals 39 61.9 18 28.6 57 90.5
~.
I
unidentified birds 3 4.8 3 4.8
vegetation 2 3.2 1 1.6 3 4.8
TOTAL
items/scat
44
1.10
40
19
1.06
63
1.09