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RAPTOR STUDIES ALONG THE PROPOSED v J
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SUSITI1A POtillRLINE CORRIDORS,OIL PIPELINE
AND IN THE YUKON AND COLVILLE RIVER REGIONS OF ALASKA
cm-mINED REPORT FOR 1975
Submitted to
u.s.Fish and Wildlife Service,
Bureau of Land Management,
'"National Park Service,
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\Arctic Institute of North America
and "1
American Museum of Natural History
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Clayton If.White .
Department of Zoology
Brigham Young University
Provo,Utah 84602
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1976
Tom J.Cade
Section of Ecology and Systematics
Langmuir Laboratory
Cornell University
Ithaca,New York 14853
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OUTLINE OF CONTENTS
Introduction
A.Acknowledgements
II.General Itinerary.
III.Reports
A.Power line Corridors -Susitna Dam Project
B.The Tanana River
C.Yukon River and Some Tributaries
D.The Pipeline -Sagavanirktok River,Old Man and Prospect
E.Colville River Watershed
Discussion and Recommendations('
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IV.
A.
B.
C.
D.
E.
The Use of Fixed-wing and Helicopter Aircraft
The "Old Man"and "Slope Mountain"Sites for Falcons
The "Hoodoos"
The "Critical Habitatll Concept and Raptors
Importance of the Colville and Yukon Watersheds OR Where Falcons
Are and Where They Are Not
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INTRODUCTION
The general purpose of our studies in 1975 was basically twofold:First,
to determine the distribution,density,and status of falcon populations in
specific areas of human activity,in order to assess and predict potential
disturbances and to make recommendations to help mollify such impacts.Second,
to survey areas of high falcon density in areas of minimal human influence,
where there is an historical record of the population and where actual or
reported declines in the populations have occurred.
Much of the area covered by our studies was viewed in relation to the
"critical habitat"concept as it pertains to both the Peregrine Falcon and the
Gyrfalcon.Also,we tried to evaluate the effectiveness of the use of aircraft
in such studies.These topics are discussed at the end of our field reports.
Projects under the first objective included:
1.A survey of the proposed power line corridor from the Susitna Dam Site
projects.This area is basically a route following the existing circular high-
way system from Anchorage to Fairbanks,plus the Denali Highway.The Dam Sites
proper were surveyed in 1974.
2.A limited check of critical localities along the Alyeska Pipeline
Route,principally along the Sagavanirktok'River.Areas near the Prospect
and Old Man Caops were also checked.
The second objective included:
1.A comprehensive survey of the Tanana River from Tetlin Junction to
Nenana.In part,this river follows the power transmission lines of the
Susitna Project (see above)over the portion from Delta Junction to Nenana.
2.An extensive survey of the Yukon-Charley River areas,principally
that portion of the Yukon between the mouth of the Fortymile River and Circle,
Alaska,and the main Charley River downriver from Crescent Creek.
3.An extensive survey of the Colville River watershed,including the
main tributaries from the base of the Brooks Range to their confluences and
the Colville River from its headwaters to Ocean Point.
These studies were supported by the U.S.Fish and Wildlife Service,the
Bureau of Land lmnagement,and the National Park Service.In addition,some of
our travel expenses were defrayed by grants-in-aid from the Arctic Institute of
North America and the Frank M.Chapman Fund of the American Museum of Natural
History.l1any persons were helpful in the accomplishment of our work.In
particular we acknowledge the assistance of Melvin Monson,Jerry Reid,Donald
Thurston,J.W.Smith,and Virgil James of the FWS:Steven Leskosky,Eugene
Ludlow,Gerald Timmons,James Lofton,Tom Sadowski,Roger Bolstad,and Skip
Ambrose of BLM:and William Brown,NPS.Harry Quinn,Texaco Oil Company,also
provided us with some transportation out of Umiat.
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June 10-12
June 12-15
June 15
June 16
June 17
June 18
June 19
June 20
June 21 -July 3
July 4-7
July 8-10
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General Itinerary
Anchorage
Power line route:from Anchorage to Talkeetna to
Cantwell to Carlo Creek,Matanuska River,Glenn
Highway,Richardson Highway to Black Rapids,Denali
Highway
Overflight of the Fortymile and Charley Rivers
Tanana River
Fairbanks
Power line route:Tanana River below Fairbanks,
Anchorage-Fairbanks Highway between Nenana and Carlo
Creek including the Nenana River
Fairbanks
Alyeska Pipeline route:Sagavanirktok River,Old Han
and Prospect Camp areas
Colville River watershed
Barrow and environs to Seward Peninsula with Secretary
of Defense Schlesinger
Yukon and Charley Rivers
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Po\-lliP.LINE CORRIDORS -SUSITNA DAN PROJECT
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In June 1974 the proposed Susitna River reservoir impoundment area,
starting at Devils Canyon and running to within ten miles of the Susitna
Glacier,was surveyed for nesting raptors,in particular the Peregrine Falcon.
Although no Peregrines were found nesting in the area (Gyrfalcons,Bald Eagles,
etc.,were found there,however),it appeared possible that falcons may nest
along the proposed corridors to transport power from the dam site to ei,ther
Anchorage or Fairbanks.Accordingly,the power line corridors as they are
currently conceived were surveyed between 12 and 15 July and on 18 July 1975.
Description of Region
The corridors generally follow adjacent to the existing highway system
and form a more or less figure-eight configuration with the middle portion of
this configuration being formed by the Denali highway.The corridors pass
through three arbitrarily defined habitat regions.
1.The cottonwood-deciduous-mixed coniferous forests (taiga)along the
Susitna-Matanuska-Copper River systems.
2.The upland areas of the Denali-Summit Lake-McKinley Park region.
3.The interior coniferous (taiga)forests of the Delta Junction-
Fairbanks-Nenana region.
These areas as thus defined are somewhat discrete units with the upland
areas being primarily occupied by Gyrfalcons,Merlins,and Golden Eagles while
the forested regions provide habitat for Bald Eagles,Peregrine Falcons,
Accipiters,and Harlan's Hawks.
1.Cottonwood-laxed Coniferous Forested P~gions
The llatanuska River Valley looks extremely good for Peregrines.
Something on the order of 35 individual cliffs occur there,but only about 10,
based on our judgment and previous experience,look right for Peregrines.
Cliffs are about equally common on either side of the river.On the downriver
portion of the river,the cliffs are generally smaller than upriver.Notably,
large cliffs occur between Sulton and Chickaloon Rivers and in the vicinity of
Sheep Mountain,with Sheep Mountain itself actually forming the last major
cliff.The portion of the Copper River along the corr~dor (Glennallen to
Gulkana Junction)is rather more like the interior forested region but is
included here for convenience.Cottonwoods still fringe the rivers,and the
cliffs are more or less high conglomerate or sandy banks,rather soft and
unstable in nature.Lastly,the Susitna River portion of this corridor flows
through generally unsuitable falcon country where only small cliffs occur
(except upriver from Devils Canyon itself,but this is outside of the corridor
and was treated in the report for 1974).Away from the corridor,the mountains
contain rather significant cliffs which might harbor Gyrfalcons (although none
were seen nor were any really outstanding falcon signs or perches found),but
they are generally too far from the corridor to.be significantly affected.
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2.Upland Areas
GenerallYt a tundra or low vegetation region.Many excellent cliffs
occur in this region,especially on the northern side of the Denali Highway,
the northern end of Summit Lake and upper Delta River region,and the entire
canyon between Yannert and Lignite.The Nenana River flows through a significant
portion of the western side of thiq region,.although the upriver portions flow
generally through flat,rather cliffless country.
3.Interior Forests
The interior forests are those areas along the corridors north of the
Alaska Range.PhysiographicallYt the region is quite distinct since the river
valleys are large and flat,the rivers wide,braided,and open with the cliffs
generally forming the northern (right)limit of the river as contrasted to the
~Iatanuska,Copper,or upper Nenana Rivers,which are rather enclosed by steep
canyons.
In all about 800 miles were covered in a survey of these regions.An
additional 80 miles of the Denali Highway were surveyed in 1974.
Methods and Itineraries
We carried out these surveys principally by air with either a fixed-wing
aircraft (Helio Courier)or helicopters (Bell Jet Ranger 206B,G-2 Bell t Boeing
BO-lOS),although the section from Eureka to Glennallen was covered by automobile.
A specific itinerary is given in Table 1.In addition to the pilot there were
always two observers (C.M.\~ite and T.J.Cade)and along the Glennallen to
Black Rapids and Denali Highway sections we were accompanied by Tom Lofton of
BLM.Our basic mode of operation was to search out cliffs along this corridor
and fly as close and as slowly by them as weather conditions and pilot's
judgment permitted.It might be mentioned here that to be successful in much
of this type of work the experience,confidence,and mental attitude of the
pilot are of extreme importance.In many cases,several passes were made by
specific cliffs usually at speeds of 30-60 mph.In the case of the Matanuska
River t checks were made both going up and coming down the river.Notes were
made on suitable cliffs or occupied cliffs.Old "eyrie-like"sites (green
grassy ledges with what appeared to be old or in some cases recent falcon
activity)were also noted.The most successful observations were made in
helicopterS t except in the case of Gyrfalcons and Golden Eagles,for which the
Helio was about equally satisfactory.
Results
Susitna Valley to Carlo
Other than a pair of Bald Eagles at Chulitna Pass,the only cliff-
nesting birds we found were Golden Eagles and Ravens.Ravens with young still
in the nest were found in East Fork.An additional Raven's nest with young was
found near Carlo.In general,the highest density of Golden Eagle nests was
between Cantwell and Carlo;four were along Carlo Creek itself.
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TABLE 1.Itinerary of travel for the Susitna Dam Power
Transmission Corridor,1975
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Date
12 June
12 June
12 June
12 June
13 June
13 June
13 June
13 June
13 June
13 June
13 June
14 June
14 June
14 June
Time
1035-1130
1148-1345
1445-1630
1645-1800
1100-1140
1200-1230
1230-1335
1335-1400
1500-1635
1650-1725
1830-2300
0620-0815
0845-0935
0935-1135
Aircraft and Pilot
Jet Ranger 206B
Jim Morgan,BLM
Jet Ranger 206B
Jim Morgan,BLM
Jet Ranger 206B
Jim Morgan,BLM
Jet Ranger 206B
Jim Morgan,BLM
Bell G-2
Garlan Edwards
Bell G-2
Gar1an Edwards
Bell G-2
Gar1an Edwards
Bell G-2
Garlan Edwards
Bell G-2
Gar1an Edwards
Bell G-2
Garlan Edwards
Automobile
Bell Jet 206B
Ed Tarsa,BLM
Bell Jet 206B
Ed Tarsa,BLM
Bell Jet 206B
Ed Tarsa,BLM
Area Surveyed
Anchorage,overland to Willow
to Talkeetna
Talkeetna,Susitna River,
through Chulitna Pass,adja-
cent to highway (Honolulu
Creek,East Fork Chulitna,
etc.)to Cantwell
Cantwell north along Nenana
River (Panorama Mt.)to Carlo
and return to Cantwell
Cantwell south along highway
to Anchorage
Anchorage to Palmer
Palmer,up Matanuska to
Chickaloon River
Chickaloon River to Sheep Mt.
Sheep Mt.,following river to
Eureka
Eureka down Matanuska River to
Palmer
Palmer to Anchorage
Anchorage via Glenn Highway
to Glennallen
Glennallen up Copper River to
Gakona,mouth of Sanford River,
refuel at Slana
Gakona River,Round Top Mt.,
Hoodoos
Summit Lake Pipeline Camp,
Upper Delta River to Bolio
Lake and return to Paxon
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TABLE I,continued
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Date
14 June
15 June
15 June
15 June
15 June
16 June
16 June
18 June
18 June
Time
1200-1236
0615-0740
0820-1000
1020-1205
1345-1435
0625-0900
0900-1315
1330-1515
1515-1625
Aircraft and Pilot
Bell Jet 206B
Ed Tarsa,BLM
Bell Jet 206B
Ed Tarsa,BIM
Bell Jet 206B
Ed Tarsa,BUI
Bell Jet 206B
Ed Tarsa,BLM
Cessna 210
J.W.Musgrove
Boeing BO-l05
Gene Parrish,BLM
Boeing Bo-l05
Gerie Parrish,BLM
Helio Courier
Al Wright
Helio Courier
Al Wright.
Area Surveyed
Paxon overlook directly to
Glennallen
Glennallen to Grizzly Lake to
Paxon
Paxon,Paxon Mt.,Fourteen-
Mile Lake,Tangle Lakes,
Glacier Lake,Paxon
Paxon,High Valley,Basalt
Lake,Maclaren River,Crazy
Notch,return overland to
Glennallen
Glennallen,Mentasta Pass to
Tanacross
Tanacross,Tetlin,down Tanana
River to Billy Creek
Billy Creek down Tanana River
to Big Delta and Fairbanks
Fairbanks,along Tanana River,
and Tanana River to Carlo
Carlo via California Creek
and Totatlinika River to
Fairbanks
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Matanuska River to Eureka
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The Natanuska River looks like ideal country for falcons.None-w~re
seen nor did we see any promising sign,although several good ledges exist.It
is hard to believe that at one time Peregrines did not nest along this river,
but there are no valid reports.There was a verbal communication (Tom Ray and
Richard Graham)that a pair nested at Sheep Mountain,and they might do so today.
They would be very hard to find,hO~lever,without working on foot because of the
size of the cliffs there.Ravens were common along the river with five active
nests being found,on average,a pair every 10 miles.Some of the nests in
between the active ones may also have fledged young in 1975.l~ith regard to
Ravens,the Matanuska is similar to the Susitna River,in its engorged portion,
where active Raven nests were found on average every 4.6 miles in 1974.It is
indeed unfortunate that the Matanuska River was not worked from a boat in the
late 1940's or 1950's when falcon populations were demographically healthy.
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Glennallen to Summit Lake
This area produced no raptors except for Bald Eagles using the Copper
River or tributaries of it.Nesting of the eagles was not established;however,
of five seen,four were not fully adult.
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Summit Lake -Denali Highway
This region is excellent for Golden Eagles and Gyrfalcons.The
Gyrfalcons persist in breeding on the Hoodoos,and the nest had at least one
young at the time of our visit.~10 Gyrfalcon sites are on the west side of
Paxon Mountain,and a lone Gyrfalcon was seen flying parallel to the Denali
Highway near there.Two inactive Gyrfalcon sites were found near Tangle Lake,
one three miles east of Long Tangle Lake and one a mile and a half ~lest of
Long Tangle Lake.A site which may have been used by Gyrfalcons was four miles
west of Basalt Lakes.Finally,~le observed an active Gyrfalcon nest with four
eggs about nine miles up the Maclaren River from the highway bridge.
Golden Eagle nests were found at seven separate localities between
Paxon and Little Clearwater Creek.The majority were more than two miles from
the Denali Highway and mainly on the north side.The general lack of active
Gyrfalcon nests may be accounted for by low ptarmigan numbers.
Upper Delta River
Very similar to the country along the Denali to the south.Golden
Eagles were found in three localities and a possible Gyrfalcon site at one.
They are rather removed and remote from the highway,however,and they should
be of little concern in regard to powerline construction.
Nenana to Carlo Along Nenana River
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Good raptor country does not really exist along the portion from
Nenana to about Ferry.Rather well-consolidated cliffs occur near Lignite,
and Raven and Golden Eagle nests were found.The cliff at Mount Fellow
contained two eagle nests.Once again,one might think of this section of
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country as good Peregrine habitat,but none has ever been reported.We saw no
likely sites and,in fact,when up close to the cliffs,they are remarkably
free of raptor sign (fecal droppings),except for Ravens.Gyrfalcons are
reported to occur there (Heinrich Springer,personal communication),but we
failed to find even likely looking nesting sites.However,the air was rough,
making it dangerous to fly a fixed-wing aircraft too close to all of the cliffs.
TANANA RIVER
We have left this region until last,since we also want to treat it
separately.For the purpose of the power transmission lines,it is sufficient
to say that no falcons currently nest between Delta and Nenana via Fairbanks
where the corridor is projected.Any lines should be kept far to the south of
the river in the flats,however,because of the potential for falcons to return
to the Tanana cliffs.A lone,deserted,but this year's Peregrine egg was
found in a former nesting site in the Birch Lake area.The entire right limit
of the river where there are cliffs should be left unmolested,however,and
several sections declared as "Critical Habitat"for the possible return of the
falcon (see discussion beyond).
Conclusions and Recommendations
Based on data gathered in 1974 and 1975,we conclude that the major portion
of environment that will be traversed by the power1ine corridor includes habitat
of min~r importance to cliff-nesting raptors,except in certain specific and
rather restricted areas.The following recommendations are offered.
1..The overall superior route would be to follow or use the existing
powerline route along the general course of the Anchorage-Fairbanks Highway.
This route generally parallels the highway on the east side to at least Rex.
From that point south,it would disturb the least cliff-nesting raptor habitat
if it were placed on the east side of the highway.Once beyond Cantwell,there
is essentially no problem with cliff-nesting birds.
2.It would seem that between Glennallen and Big Delta the clear-cut
choice would be to utilize the pipeline corridor or to parallel it.
3.From Palmer to Glennallen it appears to make little difference,but
least damage would be done by remaining close to the existing highway.
4.Should a line follow the Denali Highway,the south side of the road
should be used at least to Camp Denali.This would thus avoid nesting
Gyrfalcons,Golden Eagles,and Ravens.
5.The most critical portion of the transmission line,if constructed in
the figure-eight configuration,is that portion from Big Delta to Fairbanks and
thence to Nenana.Between Big Delta and Fairbanks lines should be placed to
the south of the river by some 1.5 to 2 miles,or else well to the north of the
highway.This placement would avoid conflict with peregrine falcon "Critical
Habitat."Power lines already exist between Fairbanks and Nenana,and they
essentially follow the Nenana Road but keep well back from the river bluffs,
't'7hich once again form "Critical Habitat"for Peregrines.
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B.THE TANANA RIVER
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Introduction
The Tanana River deserves a special discussion in this report.Not only
does the Tanana run along a portion of the proposed power transmission corridor,
but also it is a region in which a significant population of Alaskan Peregrines
has,for all intent and purpose,met its demise.The analysis of potential
problems arising from the location of the power line in proximity to the river
has been given above.It remains here to review our findings and to encapsulate
the data available on this river.
Hethods
A Boeing BO-105 helicopter was used to make one survey flight along the
river from Tit1in to Fairbanks on 16 June.Both known eyrie sites and c+iffs
where no eyries have existed were checked.A He1io Courier was then used to
check the portion of the river between Fairbanks and Nenana on 18 June.This
was the first time we had had experience with a He1io Courier and,thus,each
cliff was passed several times to determine the performance capabilities of the
aircraft in this work.Because of the nature of the cliffs and their size,we
felt that they were adequately examined to determine occupancy.
Results
Between Tetlin and Fairbanks we judge there are about 36 distinct cliffs
or groups of outcrops.Between Fairbanks and Nenana,9 more were examined for
a total of 45 cliffs.Of this number,19 separate cliffs are kno~qn to have been
used by Peregrines,but presumably two were alternate sites,and only some 17
pairs are thought to have been on the river in recent times (Table 2).Haugh
(1976)determined that some 19 pairs had been on the Tanana in earlier years.
It is probable that under pristine conditions before the turn of the century,
and probably up through the early 1900's,there were an additional two eyries
or so,based on the nature and distribution of the cliffs.It may be,then,
that as many as 20-21 pairs originally used this section of the river;we shall
never know.
In 1975,Raven nests were found on 16 of the cliffs,and at least seven
had either produced Ravens or had young still in the nest at the time of our
visit.At least four of"the active Raven nests were on the same cliffs formerly
used by Peregrines.Golden Eagle nests were on nine of the cliffs,and one
contained a chick.A Harlan's Hawk occupied at least one cliff and may have
occupied a second.A pair of Peregrines was at the cliff at the mouth of the
Robertson River (eyrie 2,Table 2).~..re could not find young,and the adults
acted as though they were not nesting.A single adult female peregrine was
found at a cliff in the Dot Lake area (between eyries 4 and 5,Table 2).We
could not determine whether she had a mate.Lastly,a single deserted egg,
laid in 1975,was on a ledge below the Little Delta River (eyrie 11,Table 2).
We do not know whether the single egg suggests that an u~ated adult female
laid it,whether a pair was there and then deserted,or whether it was laid by
a not fully adult female.
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C TABLE 2.Historical occupancy of cliffs and production of young,where
known,by Peregrines on the Tanana River between Tetlin and Nenana
Eyrie Pre-
No.1963 1963 1967 1970 1971 1972 1973 1974 1975
1 +?+
2 +?++(3)++(2)+(1)+(1) +(0)
3 +?++(3)+(3)+(2)+(1)-+(0)
4 +?++(3)+(3)+(0)+(3)
5 +?+
6 + +++(1)
7 +?+
8 +?++?????
9 +?++(4)+(3)+(3)+(3)
r----10 + + +
"'----II ++++(2)
12 ?+???????
13 +?+
14 +?+
15 +?++(4)?????
16 +?+
17 + +++
18 ++?
19 ++?
Total
young
produced ??20+9+7 8 1 0
+=pair present
..--?=eyrie not checked,may have been present
--=eyrie not occupied
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Conclusions
It is clear that a once viable population of Peregrines is now all but
gone.Haugh (1976)has made an analysis of possible reasons for this disap-
pearance.Irrespective of just why this decline occurred on the Tanana,for
example,and not on the Yukon River also,it is most important that the cliffs
themselves receive some sort of protection in the event a comeback starts.We
should not be content to believe that because the falcons are now gone,we
no longer need to consider the nesting cliffs when planning developments that
alter habitat.Some people have assumed that if a falcon tried to come back to
cliffs formerly used only to find that they were no longer suitable because of
a road,a pipeline,etc.,the falcon would just go to some other new but
undefined area to establish nesting.There are no data to support this notion.
Rather,specific cliffs have a history of use by falcons for a variety of
reasons.It is the specific cliffs that have been used,whether currently in
use or not,that need protection.They are the ones most likely to attract
falcons in a recolonization attempt.Such cliffs should receive "Critical
Habitat"status as well as a certain buffer zone about them.
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C.YillCON RIVER AND SO}ffi TRIBUTARIES
Description of Region
The area under consideration here consists of the main Yukon River from
Fortymile,Y.T.,to Circle,Alaska,and certain tributaries that drain the
uplands lying between the Yukon and Tanana Valleys,the Fortymile River itself,
the Seventymile River,the Charley River,and Birch Creek.This area lies
entirely within the Northern Plateaus Physiographic Province,which includes
a large wedge of ancient uplands,now eroded and dissected by alluvial valleys.
Elevations range from 600 feet at the border of the Yukon Flats to more than
6400 feet at the divide above the head of the Charley River.Much of the area
lies within the Yukon-Charley National Rivers proposal,and a great deal of
information on the region can be found in the u.S.Department of the Interior's
"Final Environmental Statement"on that proposal prepared by the Alaska
Planning Group.
The bedrock geology is extremely complex in this region,partly because
the Tintna Fault Zone cuts it into two different geological provinces and partly
because of a remarkable variety of exposed rocks ranging in age from Precambrian
to Tertiary.South of the fault,in a region including most of the Charley
River,the upper Seventymile,and the Fortymile,a sequence of metamorphosed
Precambrian and Palaeozoic sedimentary and volcanic rocks interfinger large
granitic bodies that were injected into the earth's crust during the Mesozoic.
North of the fault,along the main Yukon and north of it,lies a remarkably
inclusive series of unmetamorphosed sedimentary rocks varying in age from Upper
Precambrian to Upper Tertiary.Most of these rock strata are exposed as cliffs
and canyon walls along the Yukon and its tributaries.
Despite the generally wooded and forested nature of the Yukon ecosystem,
the main watercourses open up the landscape sufficiently to provide suitable
hunting habitat for falcons.In addition,fires,lakes,and bogs also con-
tribute to the maintenance of open lands suited to the Peregrine's style of
hunting.These features together with the extensive cliff-~ike formations
associated with the rivers 'combine to create good to excellent nesting habitat
for Peregrines throughout the central Yukon drainage systems both north and
south of the main river.
The river bluffs of the main Yukon have been known as prime falcon habitat
since the Gold Rush days of the late 1890's (Bishop,1900).They were first
examined in detail in 1951 (Cade,1953,1960)and have received increasing
attention by a number of investigators since 1966 (Cade,l{hite,and Haugh,1968;
Enderson,Roseneau,and Swartz,1968;Temple,Enderson,and Swartz,1970;
Enderson,Temple,and Swartz,1972).
l1ethods and Itineraries
We surveyed these rivers by fixed-wing aircraft (Cessna 210 turbo or Helio
Courier)and in a Bell Jet Ranger 206 helicopter.Table 3 shows the itinerary,
personnel,and aircraft involved.There were two or three observers in addition
to the pilot,and our method of survey was to fly along watercourses and pass
as close as possible to cliffs where falcons might be nesting.We made repeated
TABLE 3.Flying itinerary for the Yukon River and tributaries in 1975c
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Date Time
15 Jtme 1710
1730
1740
1810
1840
8 July 0715
0732
0822
0828
0921
0945
1023
1032
110S
1231
1241
1315
1340
1425
1431
1539
1552
1630
1712
1726
1735
Area Surveyed,etc.
Cadet White.and R.J.Trimble of BUM flying with J.W.
Muskgrpve.Cessna flO.at mouth of Fortym1le River.
Passing over Eagle.Village.
Head up the Seventymile River and across to mouth of Charley
aiver.
Head up Charley River.
Passing over source streams of the upper Charley.
Cadet White.and Virgil James (FWS,LE)flying with Bill
Lynch for Al Wright's Flying Service in Helio Courier,at
Yukon River 2 mi W Circle headed W to Twelvem1le Bluff.
Down at Circle for fuel.
Off upriver to survey cliffs.
At Sixm1le Bluff.
At Takoma Bluff.
At Woodchopper Volcanics.
At McGregor'~Bluff •.
At mouth of Charley.break for straight run to Eagle for fuel.
Down at Eagle.
Off upriver to mouth of Fortymile River.
At Fortymile Bluff,check and then head downriver.
At Old Man and Old Woma.n.
At Castle Rock (Pedro Dome).
At Eagle Bluff,checking.
Down at Eagle for fuel.
Off downriver to check cliffs.
At Calico Bluff.
At Montauk Bluff.
At Kathul Mountain.
At Chester Bluffs.
At mouth of Charley River.break and head for Fairbanks.
r--
"-.-/
9 July 0930
1103
Cade and William Brown (NPS)flying with Bob Barnes for Merrie
Helicopter Service.Bell Jet Ranger II,at Yukon 2 mi above
Fort Yukon.
Down at Circle for fuel.
TABLE 3,continued
"
'\
(~--~
""------
"-----
~,-
Date Time
9 July 1159
1203
1230
1236
1259
1306
1320
1341
1425
1452
1506
1539
1827
1900
1912
1951
2014
2030
2040
2112
2130
10 July 0842
0900
0916
0918
0939
1005
1015
1025
1035
Area Surveyed,etc.
Off and upriver to the Charley.
At Sixmile Bluff.
At Takoma Bluff.
At the Woodchoppers.
At McGregor'8.
At mouth of Charley River and head upstream.
Land to Check Peregri~e,between Everett and Uncompaghne Creeks.
Off and upriver.
At cliff area No.27 just below Dewey Creek,head for Central
and fuel.
Crossing Birch Creek in the flats.
Down at Central,fuel,and then off to Circle Hot Springs.
Down at Circle Hot Springs for lodgings.
Up snd off ~o cliff area No.17 on 'Charley River.
Back on the Charley snd searching cliffs.
At Peregrine'eyrie on right limit,cliff area No.22 just up
from Highland Creek..
Land by cliff area No.29,right limit,where Brown saw a
Peregrine.
Up and off again,upriver.
At cliff area No.34,left limit 2.5 mi below Crescent Circle,
Peregrines.
Head back to Central for fuel.
Down at Central.
Back at Circle Hot Springs for the night.
Off to Chester Bluffs on the Yukon.
Pass by the Woodchoppers.
'At Chester Bluffs.
At Biedermann's Bluff.
At Kathul Mountain.
At the Nation Bluff,Peregrines with young.
At Montauk Bluff.
At Sheep Creek Bluff.
At Calico Bluff.
I
'.
r"-"~---./
TABLE 3,continued
Date Time Area Surveyed,etc.
1048
1122
1334
1342
1348
1432
1509
1520('"
\~-j
"'----_..
'----,
10 July Land at Eagle for fuel.
Up and off up Mission Creek and over pass into Seventymile
River.
1145 Land at Alder Creek.
1208 Off upriver toward ~a~ley River drainage.
1220 Down Essie Creek to Charley River.
1223 Back at cliff No.34 on Charley,head upriver.
1323 Down on bar below cliff No.46 to look for Peregrine,7.5 mi
above Crescent Creek.
Off upriver again.
At upriver divide,follow east fork.
Turn back and hea~for Central.
Down at Central for fuel.
Off to Birch Creek.
On Birch Creek where it breaks into flats and follow up to
headwaters via Crooked Creek.
1703 Down at Fairbanks.
r'----
~-
~--
"'-----
"-10 -
passes by the cliffs,until we were sure we had seen as much as could be seen.
For purposes of comparison with earlier studies,we concentrated on the
main Yukon between Fortymile and Circle,where there is a history of information
going back to 1951,but we did not land to check any of these Yukon cliffs on
the ground.Fortunately,Skip Ambrose of BLM and Bob Ritchie of RRCS,Ltd.-,-
made boat trips on the river between Eagle and Circle later in July and August,
and they checked all known nesting localities on the ground.Also,Dr.John R.
Haugh traveled dO~1Uriver by boat from Fortymile to Eagle in the same time
period.These men have made their information available for this report,and
so we are able to compare the results of our aerial survey with their ground
studies.
Findings and Comparisons with Earlier Years
1.The 11ain Yukon
Table 4 summarizes our findings for Peregrines.From the air,we
found evidence of birds present in 1975 at nine cliffs,although the one at
Thanksgiving Creek appeared to us to have been abandoned by the time of our
observations.We counted a total of 10 to 12 young at five eyries and sus-
pected from the behavior of the adults that young were also present at Calico.
The ground teams were able to account for a total of 14 cliffs where Peregrines
were seen or heard,and 10 of them held breeding pairs (including Calico and
Thanksgiving Creek)that produced 16 young.Thus,our aerial survey failed to
locate all pairs and all young,even though it was conducted at abo~t the
optimum time for finding the nests,that is,when there were large do1,;vny young
present.There is no question that the forested aspect of the Yukon region
and the complex,rocky structure of these cliffs make accurate counts from the
air very difficult--far more so than is the case in the Colville River region
of the Arctic Slope.
In addition,we flew past "T1,;'lelvemile Bluff"d01,;Ynriver from Circle on
two different days in the Helio and in the helicopter.This cliff is an old
historical eyrie.Carl Dasch owns a cabin there and reported Peregrines
present at least through 1965.D.G.Roseneau found an adult female Peregrine
with three fledged young at this location in September 1974,the first time it
has been used in recent years to our knowledge.We saw no sign of use in 1975.
We also found several old Golden Eagle nests along the Yukon,but we
missed one occupied nest on the Woodchoppers,a location that we knew about
and flew right by.Recently emptied Ravens'nests were common,and we found
one occupied Harlan's Hawk nest on a cliff.
Table 5 summarizes the information we have on the occupancy of cliffs
by Peregrines between Fortymile and Circle for eight years between 1951 and
1975.On compiling this list,we were surprised to discover that no less than
31 different localities have been used by Peregrines one or more times,although
20 is the maximum number known to have been occupied in anyone year (1951).
(There are more than 40 potentially usable cliffs between Fortymile and Circle,
a distance of about 200 miles.)Only"9 cliffs have been used in most years (in
all or all but one of the recorded years),while 10 have been used only once or
c TABLE 4.Peregrine eyries and sightings in the Yukon River Region.1975
Cliff Location Date Findings
A.Main Yukon River from mouth of Fortymile River to Circle,Alaska
1.Castle Rock (Pedro Dome)
2.Calico Bluff,no.10
3.Sheep Creek,no.14
4.Montauk Bluff,no.15
5.Nation River Bluff,no.17
6.Kathul Mountain,no.22
7.Woodchopper Volcanics across'
from Webber Creek,no.29
8.Mouth of Thanksgiving Creek,
no.30
9.Twelvemile Bluff,no.35
8/1'
8/7
8/7
8/7
10/7
8/7
8/7
8/7
8/7
Adult seen but it did not defend.
Aggressive pair but nest not seen.
Pair and 1 large doWny chick.
"Adult female perched.no defense.no nest
~een.
3 or 4 large downies,partly feathered.
1 nestling seen on ledge,no adults.
Adults and 3 downy young.
Empty scrape but shelf looked used this
year.
Adult female feeding 2 or 3 young.
Charley River.from mouth app~oximately 78 miles upriver into east source
5.2.5 mi below Crescent Creek,
no.34
6.1 mi above Crescent Creek.
no.38
7.7.5 mi above Copper Creek,
no.46
(--,\B.
"-....../1.
2.
3.
4.
Between Everett and Uncompaghne 9/7
Creeks.no.3
Opposite Todd Creek.no.7 9/7
Upriver from Highland Creek,9/7
right limit no.22
5 mi below Crescent Creek.9/7
no.29
9/7
10/7
10/7
Adult female seen flying and perched on
cliff.
Green,peregrine-looking nest ledges.
Adult female with 1 young about 3 weeks
old..
Bill Brown saw an adult flying by cliff
but not seen when we landed..
Pair of adults allover cliff,no nest
seen.
Old falcon ledge with green grass.
Adult female flying over great cliff.
stooped at helicopter but no nest seen.
C.Birch Creek from edge of foothills upriver 62 miles to head of Crooked Creek
roO
...:......--~.
1.2 mi below Sheep Creek,no.12 .10/7
2.Across from Sheep Creek,10/7
no.14
Single.abandoned egg on open ledge
Adult male flew from cliff (same area
reported as a site by BLM).
/-----
~-~
.,
..
C'TABLE 5.Yukon River Cliffs used by Peregrines.Fortymile to Circle
~'
Cliff Years Total Class "A"
Number 1951 1966
1967 1965 1970 1971 1973 1975 Occupancy Eyries 1
'.
Fortymile River.Yukon Territory
1 +no +-++++6/7 X
2 -no -++--+2/7
3 ?no ?++.+",+-4/7
4 -no + +----2/7
5 ++++++ ++S/S X
6 +-------1/8
7 +-?+.+ +++6/8
8 -+-- - ---1/8
9 +-------1/8
Eagle._Alaska
(~10 ++-++-+-5/8 X
"
"'---/11 +-- - ----1/8
12 ?+ +
+.+ + + +
7/8 X
13 +------ -
1/8
14 --+--+--2/8
15 + + +-+-++5/8
16 ++++++++8/8 X
11 + ++ ++++ +
8/8 X
Nation River
18 ---- -
+-+2/8
19 ++ + +++++8/8 X
20 --- --+--1/8
21 +--+----2/8
22 +++ +--+-5/8
23 -+- -----1/8
24 ?-++++++6/8 X
25 ++++++++8/8 X
26 ?+?++-++5/8
~"
..
C')TABLE 5.continued
Cliff Years Total Class "A"
1966
,
1968 Eyries lNumber195119671970 1971 1973 1975 Occupancy
27 +++----+4/8
28 ++++ +---5/8
29 + + + + ++--6/8
30 ++ + + +++ +
8/8 X
:I'-
31 ++ ++----4/8
"
Circle,Alaska
1951 data'from Cade (1960);1966 data from Cade et al.(1968);1967 data fra.
D.G.Roseneau,L.G.Swartz,J.H.Enderson,and J.Campbell;1968 data fro.T.J.
Cade,C.H.White,and J.H.Enderson;1970 data from J.R.Enderson,S.A.Temple,
and J.Campbell;1971 data froaT.J.Carle and'L.G.Swartz;1973 data fraa C.M.
White,S.K.Sherrod.S.Ambrose,and R.Ritchie.
~',
~
-~
1 .
Based on Hickey's (1942)classification of eyrie quality.
+•site occupied by a single bird or pair
- •site unoccupied
? •site probably occupied
no ~site not checked
CJ
r-'\
"'-/
/~---
"--.-/
-11 -
~lice.Three cliffs known to have been used in 1951 have not been occupied
since,although two of them were said by the local Indians to have been occupied
"for many years"prior to 1951.
In 1951,Cade recorded Peregrines at 20 locations between Fortymile
and Circle,and 16 of these eyries had productive pairs that fledged at least
20 young.Because this was the first time Cade had been on the Yukon and
because his single survey was carried out late in the season at or just after
fledging time,these figures must be considered as minimal reflections of the
actual numbers present.With the data for more recent years to provide a
perspective,it seems likely that pairs were missed in 1951 between Fortymile
and Pedro Dome,at Calico,and perhaps at Thanksgiving Creek and between
McGregor's Bluff and Eureka Creek.Probably there were between 20 and 23 pairs
of Peregrines nesting along the main Yukon in the 1950's,producing about 30
young per year.
The data for 1966,derived from four trips up and do~~the river,do
not differ significantly from this revised estimate for the decade of the 1950's
(Cade,t~ite,and Haugh,1968).Seventeen pairs nesting between Pedro Dome and
Circle fledged 30 young,and it is probable that two or more pairs resided on
cliffs upriver between Pedro Dome and Fortymile that year,although we made no
observations on that stretch.The year 1968 was also similar:Nineteen cliffs
were occupied,at least 17 pairs attempted to nest,and a minimum of 12 and
probably as many as 16 young fledged from the eyries downriver from Eagle;the
number fledged upriver from Eagle was not determined that year.The first year
to show a reduced number of occupied cliffs was 1970 when seven productive pairs
raised 18 young (Temple,Enderson,and Swartz,1970).The population appears to
have remained essentially unchanged since then--around 14 to 15 occupied cliffs
per year with a yield of about 16 young per year from 9 or 10 productive pairs.
For example,L.G.Swartz and Cade found 9 productive pairs at the end of the
nesting season in 1971 with 15-16 young,figures that compare closely with our
1975 findings.
Of the ten cliffs judged to be most superior nesting habitat for
Peregrines--based on physical characteristics and knOwn occupancy--only one,
Eagle Bluff,has become unproductive since 1970.Five cliffs that were used
fairly regularly from Takoma Bluff downriver to Circle have been largely
unproductive since 1970,and only one superior cliff on this stretch remains
active.
2.Fortymile and Seventymile Rivers
Our examination of these watersheds consisted of quick,aerial views
of potential nesting habitat for Peregrines,and we did not do any close search-
ing for actual falcon eyries.Seen from the air,both rivers reveal good to
excellent potential for holding pairs of Peregrines.The Fortymile with its
three main branches is an extensive drainage,which is associated with large
and frequent rocky cliffs along its steep-walled valleys,and provides more
than 150 miles of possible falcon habitat in Alaska alone.The Middle Fork
begins to cut bluffs that look suitable for falcons about five miles below
Joseph,the North Fork has outcrops beginning about five miles below Inde-
pendence Creek and is particularly impressive for several miles up and downriver
..
(';
(~
"'---
'--..---
-12 -
from liThe Kink,"while the South Fork shows good habitat from Chicken downriver.
The main Fortymile is lined internittently with cliffs and outcrops all the way
to its confluence with the Yukon.
The Seventymile River is a less branching river with an essentially
U-shaped,glaciated valley,but it is lined with intermittent bluffs and rocky
outcrops all the way upstream to Diamond Fork.Most of these cliffs are low
and small by comparison with those on the Fortymile,Yukon,and Charley Rivers;
but this river,some 60 miles long to the mouth of Diamond Fork,has potential
for four or five pairs of Peregrines.Both the Seventymile and the Fortymile
Rivers should be thoroughly checked on the p,round for the presence of nesting
Peregrines.---
3.Birch Creek
Table 4 summarizes our observations in this area.From the point
where Birch Creek breaks out of the foothills into the Yukon Flats upstream to
Wolf Creek,a distance of about 30 niles,there are 20 rocky outcrops that
might be used by Peregrines for eyries.As on the Seventymile,most of these
cliffs are low--under 100 feet high--and small,but there are a few better
looking ones.Ue saw a single,abandoned Peregrine egg in a scrape on an open
ledge on a low,rocky cliff on the left limit about two miles below Sheep
Creek,and an adult male flew from a large cliff on the left limit across from
the mouth of Sheep Creek.The latter area is the same as that indicated on a
BU1 map as "raptor nests."It seems likely that the egg and the falcon belonged
to the same failed nesting attempt and that two eyries are not indicated by
these observations.No suitable nesting rock exists on Birch Creek above the
mouth of Wolf Creek.
4.The Charley River
Of all the tributaries draining into the Yukon between Dawson and
Circle,the Charley River shows the most promise as a region with significant
nesting habitat for Peregrines,and it certainly should be examined in detail
as soon as possible,for important land use decisions affecting the region will
soon be made in connection with the provisions of the Alaskan Native Land Claims
Settlement Act of 1971.Running for approximately 78 miles from its mouth up
through steep-walled valleys and canyons to its eastern headwater branch,the
Charley River has no fewer than 49 major cliffs,anyone of which could house
a pair of falcons.In fact,from the first cliff about five miles up from its
mouth to the division into east and west headwater streams,one is never out
of sight of a major rock formation.Every bend of the river has one or more
cliffs.l~ny of these cliffs are from 200 to nearly 1000 feet high and are
often arranged in a sequence of two or three tiers,much like the Yukon cliffs
around Woodchopper.Several of the tributaries--Copper Creek,Crescent Creek,
Highland Creek,and Hanna Creek--also have impressive cliffs along their courses.
The only way this river can be surveyed adequately for Peregrines is
~the ground.Even ~nth a helicopter,it would be necessary to stop at each
cliff and climb up and do,~for an hour to several hours at each place,in
order to be certain whether falcons were present or not.In the short period
of time we had available for our survey,no precise information on the number
c/
(-~"
"-_/
..........--~/
-13 -
of breeding pairs could be obtained.Probably the best way to do a thorough
census of the nesting raptors on this river would be by boat.We think it
likely that a canoe,kayak,or rubber raft could be put in place by aircraft
somewhere a few miles below the confluence of the upper tributary branches and
worked do~mstream from there.The river is shallow,fast,and rocky,with many
sharp bends,and an aluminum canoe would probably be the best craft.It might
also be possible to work this river by hiking,but it would be necessary to
cross the river frequently,and it might not be possible to find safe fording
places very often.
Again,Table 4 summarizes our findings.We saw Peregrines at four and
probably five 't·Tidely spaced cliffs,and we saw clear signs of falcons at two
other places.The pair below Crescent Creek certainly acted as though a nest
with young was present,but we could not find it.The places where we saw
falcons or sign of falcons indicate that,unlike the situation on the main Yukon
where most nests and perches are down near the river,these Charley River
Peregrines were mostly on rocks considerably up and away from the stream bed--
usually with some forest between the used rocks and the river.We think this
difference may be related to the fact that most of the open country suitable
for falcon hunting along the Charley is in the uplands at or above timberline,
while the river valley itself is quite narrow and enclosed~until it reaches
above Copper Creek,where the surrounding country begins to open up.By nesting
and perching high up near the top of the valley walls,the falcons are conven-
iently near good hunting areas and do not have to climb up and out of the river
valley to find their food.
Considering the fact that our hasty survey by helicopter yielded
sightings of Peregrines at four or five places on the Charley River,we would
not be surprised if a ground survey turned up eigh~or ten occupied cliffs with
perhaps four or five productive pairs,or possibly more if some of the larger
creeks are also included.If these numbers actually obtain,then the Charley
River would take on a significance second only to the main Yukon and Porcupine
River as a critical refugium for the nesting of taiga Peregrines.
The area is also rich in old Golden Eagle nests and P~vens'nests.
One nest of Harlan's Hawk was also found on a bluff near the mouth of the
Charley River.
5.Yukon River -Stevens Village to Tanana
The region described here covers a distance of about 147 miles between
the old abandoned town site of Fort Hamlin (downriver a few miles from Stevens
Village)and Tanana.The last cliffs actually end upriver from Tanana about
15 miles,so the area of importance extends over about 125 miles (Fort Hamlin
to 12 miles upriver from Tanana).This region was surveyed in 1974 by C.M.
White and S.K.Sherrod and again in 1975 by Skip Ambrose and Bob Ritchie.
Although the region was not checked as a part of our overall 1975 survey the
data bear presentation.
Other than casual observations by interested parties (ltl11ard Heiner,
USGS,traveled the river in early June of 1972 and told us "every suitable
cliff between the pipeline crossing and Tanana had fa1cons ll)no specific studies
(~
"--
r'
~..
'''"---~-~
-14 -
have been done on this portion of the Yukon prior to the 1974 survey.We
therefore have no historical base of comparison.At least eight different
cliffs have been used by falcons,but this occupancy possibly represents only
seven pairs.The most we have found in any year was six pairs.Based on the
six known pairs the density of falcons is about one pair per 20.8 miles of
river.The density is thus about one-fourth less than that of the Yukon between
Eagle and Circle.In addition to the cliffs with falcons there are at least
four other cliffs that we judge to be good for falcons,of which the most
striking ekamples are the cliffs on Raven Ridge and at Point No-Point.In
former years perhaps ten pairs occupied this segment of river in which case
the density would have been about one pair per 12 miles,a value more in line
with other findings on the Yukon.The 1974 and 1975 data are presented in
Table 4.
Introduction
(')D.
-15 -
THE PIPELINE -SAGAVANIRKTOK RIVER,OLD HAN AND PROSPECT
r~,
~...
'---
Since 1970,there have been biannual surveys for raptors along all or some
portions of the Alaska Pipeline.In 1970,the entire route was covered,and it
was clear that the portion north of the Yukon River was of main concern.In
1972,about two-thirds of the route was covered,but the concentrated effort
occurred north of Fairbanks and along the Tanana River.In 1974,only that
portion north of Fairbanks and the Tanana River were surveyed with most effort
north of the Yukon River.On 20 June 1975,we looked at portions of the
Sagavanirktok River and areas around two camps,primarily for Peregrines and
Gyrfalcons.Reports for the three previous years are to be found in the
Anchorage Office,U.S.Fish and Wildlife Service.
Hethods
A fixed-wing Helio Courier was used to check cliffs.Standard methods of
flying by the cliffs as slowly and closely as possible,making several passes,
were used.No ground checks were made because of difficulty of finding
adequate landing places.The cliffs adjacent to the Sagwon Airstrip were not
checked owing to the flight restrictions on the area imposed by BI1!.
Results
In past surveys,as many as three pairs of Peregrines have nested on
Franklin Bluff,but apparently only OjO pairs have been there on average since
1970 (Table 6).Usually only one of the pairs has been successful in fledging
young in anyone year.In 1974,the upriver pair near the Bruce bench mark
could not be found.He did not have adequate looks at the cliff,how'ever.lve
likewise ,\:jere unable to get on the ground to examine the area.The lower pair
about two miles north of the Greta bench mark was nesting in the usual site
and had either eggs or young,but we did not flush her.
At Sagwon Bluffs we had to overfly at such a distance that we could
determine nothing.Harry Reynolds,Alaska Department of Fish and Game,walked
to the area in July and found two pairs in the localities opposite the Sagwon
Airstrip that they occupied in 1974 (personal communication).He had not
determined productivity at the time of his report,however.From what could
be gathered,it appears likely that three pairs were present on the
Sagavanirktok River.
Two Gyrfalcon sites upriver from Sa~ron were occupied,one with four
young and one ~nth two young.Additionally,in spite of the gravel-obtaining
activity on Slope Uountain,Gyrfalcons \-Tere occupying a nest with two young
less than one-half mile distant.The Rough-legged Hawks that had nested above
the gravel pit in 1974 were not there in 1975..
The Gyrfalcon eyries to the southwest of Prospect Camp and to the north
of Old }mn Camp were checked.The former is of interest because it occurs in
a taiga situation and has only been occupied in one of the four survey years,
and the latter site is of interest because it is located within one-half mile
'-.'0 -16 -
c)
r~
"-~>
'-..-.--
of the pipeline road.The site near Prospect Camp was unoccupied in 1975.
The site south of Old Man was not used in 1970,an apparent attempt occurred
in 1972,young were fledged in 1974,but it was not in use in 1975.
Conclusions and Recommendations
1.It appears that the Peregrine Falcon population along the Sagavanirktok
River has stabilized since 1970 and is holding its own.Reproduction,however,
may be lower than desirable.Because of the lack of precise data from earlier
years,it is hard to IOlOW just how many falcons originally used Franklin Bluffs.
John Koranda,Lawrence Livermore Laboratory (personal communication,September
1975),has supplied data from 1958.In tllat year five pairs nested on Franklin
Bluff,four on the bluff proper,and one on an outlier of the bluff to the north
of the main section (see Table 6).~~o of the pairs there today are occupying
the same sites they did in 1958.Additionally,Koranda found two pairs on the
~fuite Hills,Toolik River,to the west of Franklin Bluffs,in 1958 and one pair
only in 1973.
These data suggest that only about half as many birds nest there today
as in former years.Unfortunately,we do not have comparable data for the
Sagwon Bluffs except to know that Peregrines occupied the site opposite the
building at the airstrip in the late 1950's (Marvin Mangus,personal communi-
cation)and in 1964 (David Anderson,personal communication).
2.The Gyrfalcons along the Sagavanirktok River seem to vary some"l7hat
but apparently in direct relation to food supply.They appear to be holding
up well.
3.The presence and nesting success of both Gyrfalcons and Rough-legged
Hawks using the east side of Slope Mountain and the Gyrfalcons south of Old Man
Camp should be carefully monitored.Both situations will provide an indication
as to the amount of disturbance these two raptors will tolerate.
()
'"/"---,/
TABLE 6.Sequence of utilization of cliffs bi Peregrine Falcons
on the Sagavanirktok River
Area
Franklin Bluff Section
Localitios
1958 1970
Years
1972 1973 1974 1975
1.Outlier
2.Mud site
3.Greta north
nesting
1
nesting
?
nesting
??
?
??
('\
"'---,
4.Greta
Site A
_ite B
5.Bruce
Site A
Site B
Site C
nesting nesting nesting deserted nesting
nesting
nesting
nesting nesting nesting
nesting
6.South Bruce
Sagwon Bluff
Localities
1.Red Rock
adults
present
?
?
lone Gyrfalcons
adult
1
?
lone
adult
Gyrfalcons Gyrfalcons
2.Cut
3.South
1
?
??
?
?
nesting
nesting
present
present
/'
"-_/
1958 and 1973 data are from John Koranda.
- -cliff examined,falcons not present
? -cliff not examined,falcons may have been present
r'"'---~
~~'
~"
-17 -
E.COLVILLE RIVER WATERSHED
Description of Region
The Colville River is the major drainage system of the Arctic Slope with
a watershed of about 23,000 square miles,as determined from Raisz's Landform
Map of Alaska,1948,and cliff-lined watercourses totalling more than 2000 miles
(Figure 1).Although its principal sources lie in the Brooks Range above 3000
feet and its lower reach runs through.the low-Iyin~Arctic Coastal Plain to
Harrison Bay,most of the main river and most of its tributaries flow through
rolling foothills and upland terrain varying from about 500 to 2500 feet in
elevation.Various Cretaceous rocks underlie these foothills--the Torok
Formation,marine sediments of mostly shale ~dth varying amounts of conglomerate,
sandstone and siltstone,and the Nanushak and Colville Groups,intertongued
deposits of marine and non-marine sediments consisting largely of sandstone and
clay shale,with considerable local deposits of coal,bentonite,and tuff.
Below the Etivluk,these groups overlie the deeper and older Torok Formation
and are folded to form synclines and anticlines.Variations in the structure
of the formations within these groups result in persistent east-west ridges,
mesas,and hills.
The exposed rock formations and the watercourses combine to create optimum
nesting habitat for cliff-nesting species throughout most of the Colville water-
shed,and suitable nesting habitat for falcons,hawks,eagles,and ravens is
absent only from the lower Colville below Ocean Point and along all but the
upper reach of the Itkillik.The riverine bluffs and cliffs of the main Colville
and its tributaries have been kno~vn as prime nesting habitat for Peregrines,
Gyrfalcons,Rough-legged Hawks,and Ravens since the early 1950's (see Kessel
and Cade,1958;Cade,1960;White,1964;ffuite and Cade,1971).Only in recent
years have we been able to fit the Golden Eagle into the overall pattern of
occupancy of cliff-like formations in Arctic Alaska.
Methods and Itineraries
We carried out these surveys by air,either from a fixed-~dng aircraft
(Helio Courier)or helicopter (Alouette II or Bell Jet Ranger 206).Table 7
lists the dates,times,places,and modes of travel for these surveys.Usually
there were three observers in addition to the pilot,C.M.White,T.J.Cade,
and J.W.Smith,the latter from the Law Enforcement Division of the U.S.Fish
and Wildlife Service.Our basic mode of operation was to fly along watercourses
and to pass as close and as slowly by the cliffs and bluffs as conditions,the
aircraft,and the pilot would allow.The helicopters,of course,allowed for the
slowest and closest approaches,and also made it possible to land for a ground
search,if necessary.With the Helio Courier we usually could pass within 50
feet of the cliffs at air speeds of 50 to 60 mph.In actual practice,because
of the speed,better observation was achieved at distances of 100 or more feet.
We made repeated passes by the cliffs,sometimes as many as six or eight,at
both high and low elev~tions with respect to the cliff face until we were
satisfied that we had seen all that could be seen.
For the purpose of making comparisons with previous studies,we concentrated
on the main river from the mouth of the Etivluk to Ocean Point,where there is a
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ARCTiC COASTAL "LAIN PROVINCE
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AJIlCTIC FOOTHIL.L.PROVINCE
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COLYILLE RIVER DRAINAGE
ALASKA
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Figure 1.Colville River drainage,Arctic Slope,northern Alaska.
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TABLE 7.Flying itineraries for Colville River in 1975
1735-2205 'Helio C
Main
1819-2157 He1io C
Main
1022-1450 He1io C
Main
~~,
"'----
"--._/
Date
21 June
21 June
21 June
22 June
23 June
24 June
24 June
25 June
26 June
27 June
27 June
28 June
28 June
29 June
29 June
Time
1138-1205
1433-1500
1835-1940
1040-1912
1122-1225
1005-1554
1902-2105
0935-1605
1456-1944
1015-1428
1616-1916
1036-1450
Aircraft and Pilot
He1io C
Shanahan
He1io C
Shanahan
Helio C
Shanahan
He1io C
Shanahan
He1io C
Main
He1io C
Main
Helio C
Main
Mouette II
Bidenchini
Alouette'II
Bidenchini
Alouette II
Bidenchini
Belio C
Main
Helio C
Main
Area Surveyed
Umiat upriver to High Bluff,no.52,
and return by Coal Seam,no.55
Upper Itkillik and Itkillik Lake to
head of Koyukuk.
Upper reaches of Chandler,ChancIer
Lake,down to Grandstand,over to
Umiat.
Umiat up Colville to cliff no.24,
below m.Kurupa and return,with
landings.
Umiat upriver but stormed out,return.
Umiat to no.24 and up to no.1 on
'Ettvuluk and return with stopa.
Umiat to no.52,stop for ground
BeJlrch,and return.
Umiat downriver to Ocean Point and
back with stops.
Umiat downriver to Kogosukruk mouth,
return up Kogosukruk to Umiat with
stops.
Umiat to Chandler and Siksikpuk Rivera
and re~urn,with stops.
Umiat to lover Chandler,up An'aktuvuk
into upper Kanayut and return down
Anaktuyuk to Umiat.
Umiat to Colville above Etivluk,
including lower Awuna to Section
Creek,to mouth Kiligwa and back to
Umiat.
Back to upper Colville to sources
around Thunder Creek and Meat Mountain,
to Liberator Lake,down Kttna to·Colville
and return.
Umiat to Nanushak and tributaries,
Cobblestone Creek,Peregrine Creek,
May Creek and return.
Umiat to Etivluk,up main river and
tributaries,Fay Creek,Nigu,Kutchaurak
Creek,'Otuk Creek,Iteriak Creek,return
over Kurupa and Oolamnagavik.
c\TABLE 7,continued
r
~.
/-
'-----~
Date
30 June
30 June
30 June
1 July
2 July
Time
1002-1429
1638-2020
2105-0023
1521-1830
1103-1552
Aircraft and Pilot
Helio C
Main
Helio C
Main
Bell Jet Ranger 206
(Texaco)
Helio C
Main
He1io C
Main
Area Surveyed
Umiat to East Fork of Etivluk and
tributaries,over into Kurupa and
tributaries and return.
Umiat to Oolamnagavik and tribu-
taries,l~~er Killik,Okokmilaga and
Okpikruak Rivers and return along ma;
Colville.
Umi~t to Killik upstream to AkmAlik
Creek and return via SilalinigUD
Creek and Okpikruak River and over
tundra to Umiat.
Umiat to Okokm1laga to upper Killik,
Mary Creek,Easter Creek,upper tribt
tar1el of Okokm1laga,Coal Creek and
return to Umiat.
Umiat up Colville to Driftwood and
upper Utukok,land,eogine trouble,
return.
....,,,
-18 -
c:)
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"------."
history of information going back to 1952 (Cade,1960);and we got down on the
ground as much as possible in this area,at a total of 20 locations where
Peregrines are known to have nested in the past.At no place where we made
ground observations,did we find Peregrines where they had not first.been seen
from the air.Most of our observations in other parts of the watershed were
from the air,with occasional landings where pORRible to check on PeregLines.
Table 8 summarizes our coverage of the watercourses.In addition,we flew
across the middle and upper reaches of all the tributaries from Thunder Creek
to the Killik and checked likely looking cliffs along the route.The only
portion of the watershed that did not receive attention was along the middle
and lower reaches of the Itkillik,where we ~lew from previous flights that
suitable bluffs or cliffs do not occur,and the headwaters of tributaries in
the Brooks Range,where again Peregrines are not known to nest and where flying
these sorts of surveys becomes extremely hazardous.We no doubt missed some
eagles and a few Gyrfalcons in these mountainous parts.
Findings and Comparisons with Earlier Years
1.The Peregrine Falcon
Table 9 summarizes our observations.We found Peregrines at 12,
possibly 13,sites on the Colville from the Etivluk to Ocean Point.Only three
of these sites were located upriver from Umiat.Eggs were found at nine sites
and were possibly present at one other,cliff no.7.of the 34 eggs found,we
know that seven were bad or abandoned during our period of observation,leaving
a total of 27 possibly viable eggs.If half of these eggs resulted in fledged
young--about the most optimistic expectation one could have--then 13 to 14 young
could have been produced by this population in 1975.If survival of these eggs
and young followed the pattern observed by J.Haugh in 1973 (see below),a more
likely prospect,then fewer than 10 young fledged this year.
In 1973,J.R.Haugh made a complete on-the-ground survey of the
Colville Peregrines for the.Alaska Department of Fish and Game.In June,he
found 14 pairs and a single adult between the Etivluk and Ocean Point;11 pairs
had 32 eggs.In late July,only four of these pairs had a total of nine
advanced nestlings.As in our 1975 survey,he found only three pairs at cliffs
upriver from Umiat,but they were on three different cliffs than the ones
occupied this year.Formerly,about 18 to 20 pairs nested upriver from Umiat,
while 15 to 18 nested below.(A total of 25 nesting places is known upriver,
and 23 downriver.)It is puzzling to find that the upriver nesters,which
always appeared to be most secure and generally occupied the larger,hard rock
cliffs,are now virtually gone,while the downriver falcons,nesting mostly on
low earth and mud bluffs with frequent surface slides and other forms of erosion
where egg and nestling losses have always been high,have decreased the least.
Perhaps the attractiveness of these d~mriver sites relates more to food supply
and favorable hunting conditions in the arctic Coastal Plain than to the physical
features of the actual formations used for nesting.
In 1974,C.M.t~ite made a quick helicopter survey from Ocean Point
to Umiat on 8 July at hatching time.He found Peregrines at eight places,
including five pairs with a total of 11 eggs or newly hatched young.The other
Cj TABLE 8.Miles of river covered by aerial surveys in 1975
c'
Locations
1.Main Colville,Etivluk to Ocean Point
2.Upper Colville above the Etivluk
3.Upper Itkillik River
4.Anaktuvuk and Nanushak Rivers
s.Chandler and Siksikpuk R1 vers
6.Killik River and tributaries
7.Oolamnagavik and tributaries
8.Kurupa and tributaries
9.Edvluk and tributaries .
10.Lower Ip~avik River
11.Lower Kuna River
12.Low~r Awuna River
13.Lower Kogosukruk River
14.Lower Kikiakrorak River
TOTAL MILES
No.River Miles·.
188
148
43
243
124
248
72
119
143
20
16
30
41
33
1468
r"
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•Determined from USGS Alaska Topographic Series,1:250,000
()
~TABLE 9.Peregrine eyries and sightings in the Colville watershed,1975
Cliff Location Date Findings
A.Main Colville,Etivluk to Ocean Point
~\
\~'----_/
~~~
.....-----~_.
1.2 mi upriver from Umiat,
no.55
2.Mouth of Kutcher Creek,
no.7
3.Etivluk,3.6 m1 up from
Colville.no.1
4.Upriver and Shivugak,no.59
5.KiddIe Shivugak.no.59
6.Near lIOuth of Ingaluat Creek,
no.69
7.1.0 m1 below mouth of
Kog08ukruk Il1ver,no.73
8.Mouth of lCikiakrorak River,
no.7S
9.1.5 m1 below Kildakrorak.
no.76
10.Ca.6 mi above Ocean Point.
no.79
11.Ca.3 m1 above Ocean Point,
no.80
12.U1uksurak Bluff,no.61
13.1 a1 above Dee Bench Mark,
no.67
B.Upper Colville and Tributaries
1.Tuluvak Bluff,Chandler River
2.0.5 m1.up from mouth of
Kanayut River
3.Shale Wall Bluff.Nanushak
River
4.Etivluk.ca.I m1.above East
Fork mouth
22/6
26/6
24/6
24/6
29/6
30/6
25/6
2$/6
25/6
25/6
25/6
25/6
25/6
25/6
26/6
26/6
27/6
27/6
29/6
29/6
Adult male flew off cliff.
Adult female on n~Rt.3 eggs.
Aggressive pair,no nest could be found
by ground search.
Adult female on 3 eggs,no~!:~seen.
Female not on eggs,no adult seen.
Eggs abandoned.
Pair with 4 eggs.one broken in.
Adult (female?)flies back and forth
along cliff.
Adult female on nest with 4 eggs.
Pair with 4 eggs.plus second adult
female.
Fresh scrapes;perhaps belonging to
female above?
Pair with 4 eggs.female incubating.
Pair with 4 eggs,female incubating.
Pair with 4 egp.
Adult (female?)flying silently around
cliff.
Adult female with 4 eggs,3 sloshy and
rotten.
Single adult,no screaming.
Pair,female on stick nest,at least 2
eggs.(Female would not move off neat.)
Adult seen perched on cliff.
Adult male on nest,4 eggs.
~\TABLE 9,continued
Cliff Location
5.Colville 2 m.i above mouth
of Ipnavik
6.Colville 3.6 mi (straight
line)above Kiligwa
7.Colville 3 mi below Mitten
Creek
r-"
~/.
,---.
Date
28/6
28/6
28/6
Findings
Pair with 4 eggs.
Pair with 4 eggs.
Pair,nest not seen but probable.
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-19 -
three pairs acted as though they had no occupied nests.l~ite felt that he
might have missed one or two pairs,as the survey was done hurriedly.Two of
the cliffs where he found pairs in 1974 were not occupied in 1975,and we found
falcons at three cliffs where he saw none in 1974.With the reduced Peregrine
population now in the Colville region,the pairs evidently do a considerable
amount of shifting around from year to year among the available eyries.
~fuen the information for 1974 and 1975 is compared with the earlier
data summarized in Table 10 from Peakall et a1.(1975),it appears that the
decline which began in the Colville population in 1969 may have bottomed out
in 1973 or at least that the rate of decline has lessened,for our figures on
nuWber of pairs and reproductive performance in 1974 and 1975 do not differ
markedly from Haugh's findings in 1973.~fuile one could be encouraged to hope
that the main Colville population has now leveled off at about 35 per cent of
its former breeding numbers,caution must be exercised,because the low repro-
ductive yield of less than one young per pair is not sufficient to maintain
breeding numbers without recruitment from outside sources.Possibly falcons
from the more marginal territories along the tributaries and upper Colville
move in to occupy vacancies in the better nesting habitat along the main river.
If so,the apparent leveling off or slowing of the decline in the breeding pairs
of the main Colville is deceptive for the regional population as a whole.
Throughout the rest of the Colville watershed in 1975 we were able to
find Peregrines at only seven places.Four certain nests yielded 14 to 16 eggs,
and a fifth probable nest could have raised the total to 18 or 20 eggs.Only
single adults were seen at the other two sites.Since we were working mostly
over unfamiliar ground in these areas,where we had little prior knowledge of
exact nesting locations,and especially since we conducted these surveys near
the end of the incubation period for Peregrines when the birds sit tightly on
their nests,we feel there is a greater chance that we missed some Peregrines
in these areas than on the main Colville where we could pinpoint exact locations
on which to concentrate our attention.Even so,we doubt that we could have
missed more than half of the pairs actually present on the tributaries and
~pper Col~lle.
Seven to 14 pairs widely scattered along more than 1200 miles of
watercourses in an area of some 20,000 square miles make up a very sparse popu-
lation,indeed,and lead to speculation about changes in numbers in these parts
of the Colville watershed.Have the birds always been more sparsely distributed
along the tributaries than on the main river,or have these populations under-
gone an even more severe decline in numbers than the breeders along the Colville
proper?The fact that a higher percentage of pairs has been lost on the Colville
upriver from Umiat than downriver suggests a trend that gives credence to the
latter possibility.
Fortunately,there is a little historical information on some of these
areas that provides us with insight.In 1952,along 85 miles of the Colville
upriver from the mouth of the Etivluk,T.J.Cade and G.B.Schaller found eight
pairs of Peregrines during the late nestling and early fledgling period at the
end of July and in early August (Kessel and Cade,1958).In 1975,we could find
only two pairs along the same stretch of river,and Haugh and Conner found none
during a flight on 22 July 1971.In 1956,along 35 miles of the Oolamnagavik
r
C TABLE 10.Number of breeding pairs of Pere~rine8 and productivity
on the Colville River
Total No.No.of Pairs No.of Young
Year Pairs Producing Young Produced
1952-1959 32-36 20-25 40-50
1967 27 18 34
1968 32 16 34
1969 33 13 26
1971 25 9 14
1973 14 -4 9
*Data from Peakal1,Cade,-White,and Haugh,1915
C~
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-20 -
River T.J.Cade and H.M.Webster,Jr.,found five pairs of Peregrines during
the hatching period between 30 June and 6 July (Kessel and Cade,1958),where
we could find none present in 1975.In 1968,L.L.Boyd and E.Stauber found
two pairs of Peregrines nesting on the middle reach of the Killik River,and
C.M.vlliite found two pairs on the lower Killik in 1964.J.Haugh could find
none along the entire Killik in 1971,and we could find none in 1975.In 1971,
J.R.Haugh carried out aerial surveys for the Alaska Department of Fish and
Game along the Chandler and Siksikpuk Rivers and along the Anaktuvuk and Nanushak
Rivers,where w~also worked in 1975.He found six pairs of Peregrines (also two
on the Chandler in 1972),where we found birds at only three places,single
adults at two and a pair with eggs at a third.Two of our locations were ones
at which Haugh had found Peregrines.Haugh also surveyed the Killik and Okpikruak
Rivers in 1971 and reported one pair of Peregrines on the latter at its con-
fluence with Verdant Creek and the rllddle Fork.We found none there or else-
where on these or other tributaries of the Killik system in 1975.
All of this information indicate~that (a)Peregrines were formerly
widely distributed and common on the upper Colville and on the tributaries and
(b)that the breeding populations are now greatly reduced along these water-
courses.Cade (1960)estimated that the breeding population of the Colville
drainage in the 1950's was between 60 and 150 pairs,with 100 pairs being the
best estimate of the real population.At that time he had detailed,on-the-
ground knowledge only of the main Colville,one late-summer run of the upper
Colville,and one mid-season run on the Oolamnagavik,although he had made
flights over most of the watershed at one time or another and had a good concept
of the overall topography and general ecological conditions of the entire region.
The more recent surveys of the late 1960's and 1970's confirm the existence of
suitable nesting cliffs and breeding habitat for Peregrines along all the major
tributaries except the Itkillik,and we still feel that 100 to 120 pairs is a
good,conservative estimate of the breeding population that existed in the
1950's and earlier.Conceivably it could have been even larger--around 200
pairs-as an average of one pair per ten miles of river is not unreasonable
based on data for the Colville and Oolamnagavik Rivers (45 pairs along about
302 miles of river,or about 7 miles per pair).Also,in the 1950's Peregrines
outnumbered Gyrfalcons about 3 to 1 to 4 to 1.In 1975,we located 40 pairs of
Gyrfalcons (see below),a number which again fits a concept of 120 to 160 former
pairs of Peregrines,assuming that Gyrfalcons have not substantially increased
in numbers regionwide since the 1950's.
If these speculations are approximately correct,it would appear that
the reduction in numbers has been greatest for nesting Peregrines on the tribu-
taries,next most telling on the upper Colville,and least on the lower Colville,
for an overall reduction to about 25 per cent of the breeding population present
in the 1950's.The extent to which recruitment into the breeding population
along the main Colville has been influenced by movement of individual adults
from territories along the tributaries into better habitat (probably better
feeding areas)is unknown but may have been considerable,as the local production
of young along the Colville since 1969 has been insufficient to account for the
apparent stability of the breeding population,at a reduced level,since 1973.
Such a process involving individual adults shifting from marginal territories
into more optimal range conceivably could have been going on since the late
1950's or early 1960's.
CI
('
~~
/-
"'--~.
-21 -
2.The Gyrfalcon
Table 11 summarizes our observations at 40 places where Gyrfalcons
were found.Along the main Colville 13 pairs occupied cliffs from a point a
few miles above the A,~a to Shivugak Bluff downstream from Umiat.This is
one more than the previous maximum of 12 pairs recorded in 1959 (Cade,1960)
and in 1971 Oihite and Cade,1971).By late June they had produced at least
32 advanced nestlings (actually counted)and probably 36,for an average of
about three young per pair.Twenty-five other pairs in the Colville watershed
had a total of at least 62 young,probably 68 to 70.
These figures compare favorably with our findings for recent years
along the Colville:In 1967,11 pairs had 37 young in the nest early in the
season,and they fledged 30,an average of 2.73 young per pair;in 1968,11
pairs had 28 young in the nest early in the season,and 26 presumably reached
flying age,an average of 2.36 young per pair;in 1969,9 pairs had 26 young
in early season,and all presumably fledged for an average of 2.89 young per
pair;in 1971,12 pairs produced an estimated total of 33 eggs or downy young,
but they fledged only 18 young,or an average of 1.80 per pair.
These levels of production were sustained despite generally low numbers
of ptarmigan throughout the Colville River Valley in those years ~fuite and Cade,
1971).The low productivity in 1971 was associated with an abnormally late
nesting season for many pairs,as eggs and downy young were found in late June
when the nestlings are usually 4 to 5 weeks old.Presumably weather or feeding
conditions early in the breeding season influenced this late nesting schedule.
Ptarmigan were also extremely scarce in the Colville watershed in 1975.
We made a special effort to count all birds that could be seen on our aerial
surveys.The total count was only 11 ptarmigan,all Lagopus lagopus.They
appeared to be more common along the Sagavanirktok River,where we saw six
evenly spaced males in the vicinity of Franklin Bluffs on 21 June.We '-Tere
unable to observe what the Gyrfalcons were feeding on in 1975,but ''1e do know
that microtine rodents,especially Lemmus trimucronatus and Microtus oeconomus,
were in high numbers throughout most of the region,and this rodent explosion
was reflected by a large number of rodent-eating birds in the area.The numbers
of Rough-legged Hawks were up very considerably (see below),Short-eared ~1ls
were widespread and common throughout the foothills,as well as jaegers (mainly
Long-tailed and Parasitic,but also an unusual number of Pomarine).The Jaegers
were mostly in non-breeding aggregates of 6 to 20 birds along the rivers.There
were even scattered non-breeding Snowy ~7ls between the Colville and the Brooks
Range--far south of their usual summer range on the Slope--and also a few Marsh
Harriers.It was obviously a big microtine year in the foothills,and we sus-
pect that the Gyrfalcons were feeding,as they did in 1959 (Cade,1960),on the
rodents and,especially,on the rodent-eating birds,principally Long-tailed and
Parasitic jaegers and Short-eared Owls.Certainly it would have been difficult
for them to have produced so well on the available ptarmigan.
Our data indicate that Gyrfalcons continue to thrive in Arctic Alaska,
despite considerable fluctuations in breeding success from year to year,and
there begins to be a suggestion that Gyrfalcons may actually be increasing in
numbers and shifting their ecological niche somewhat as the Peregrine population
c~TABLE 11.Gyrfalcon eyries and sightings in the Colville watershed,1975
Cliff Location Date Findings
A.Main Colville,Etivluk to Ocean Point
L 13 m1 upriver from Umiat,
no.52
2.Colville at mouth of Fossil
Creek,no.53
3.Colville near mouth of
Ninuluk Creek,no.49
4.Upriver end of Ninuluk Bluff,
no.47
21/6
21/6
22/6
22/6
Pair with 4 young,4 weeks old.
2 or more young in nest,ca.1-2 weeks
old.
Adult flying,1 or more young in nest.
Pair with 3 young,3 weeks old.
11.
12.
13.
White adult,3 young,5 weeks old.
2 young,1-2 weeks old.
3 young,3-4 weeks old.
4 young,4 weeks old.
3-4 young,3 weeks old.
Pair but nest not seen.
Pair with 3 young,3 weeks old •.
Adult,nest with 1 or more young,1-2
weeks old.
White adult,3 young,3 weeks old.
24/6
24/6
24/6
26/6
25/6
26/6
22/6
22/6
22/6
S.'Puivlik Bluff
6.1 ai downriver from mouth of
Oolamnagavik,no.41
7.Near mouth of Aupuk Creek,
upriver,no.29
8.2 mi above mouth of Kurupa
River,no.20
TSS,RlSW,Killik River
Quadrangle,no.12
10.5 mi downriver froa mouth of
Etivluk,no.6
Middle Shivugak,no.59
Umiat MOQn~ain,no.57
Tattitgak Bluff,no.54
(~
~//9.
B.Upper Colville and Tributaries
1.Chandler below mouth of
Ktruktagiak River
2.Chandler,5 1I.i below Taktu
3.Chandler below Patmagaktuk
4.Chandler,up Outpost Creek
S.Chandler,3 mi below Kutchik,
creek on right limit
6.Anaktuvuk,Schrader Bluff
7.Anaktuvuk,9 ai below Terrace
Creek
8.Anaktuvuk at Table Top
27/6
27/6
27/6
27/6
27/6
27/6
27/6
27/6
4 young,6+weeks old.
2-3 young,hard to see.
2 young,5 weeks old.
3 young,3+weeks old.
3 young.3-4 weeks old.
3 young,4-5 weeks old.
2 young,5 weeks old.
4 young,5+weeks old.
,---..,
•
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-.........._~,;.-
TABLE 11,continued
Cliff Location
9.Kanayut River
10.Colville opposite mouth of
Kuna River
Date
"27/6
28/6
Findings
2 young,4-5 weeks old.
2 young,6+weeks old.
..
o
('\
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~,
"'--/
-22 -
has declined.This latter hypothesis is suggested by the fact that in 1975 we
found two pairs of Gyrfalcons nesting not only on the same cliffs that had been
used by Peregrines but on the exact same sites on the cliffs.This would not
have been remarkable,perhaps,except that these two sites are exposed points
at conspicuous breaks in slope on completely open cliff faces--extremely atypical
locations for Gyrfalcons,which usually nest in well-sheltered sites under over-
hanging rock.(In faat,typical Oyr sites exist on bolh cliffs.)It may also
be reflected in the fact that Gyrfalcons breeding in the Colville region since
the decade of the 1950's seem less dependent upon ptarmigan for a food supply
than in earlier years when they had to exist in the same region with a Peregrine
population three to four times mora numerous than they.
In addition to the falcons,we kept notes on the nesting locations of
Golden Eagles,Rough-legged Hawks,and Ravens.Old eagle and Raven nests are
frequently used by Gyrfalcons,less often by Peregrines,while Peregrines
frequently use old Rough-leg nests and Gyrfalcons occasionally do.The nests
of these species therefore have significance for the breeding distribution of
the two falcons,as well as providing important biological information on these
species themselves..
3.The Golden Eagle
l~ile Golden Eagles are widely dispersed in sparse numbers over the
entire Arctic Slope in summer,even to the coast,they breed only in the rock-
walled valleys of the Brooks Range proper and sparingly on some of the outlying,
northern mesas and tablelands in the foothills.In 1975,we found five active
nests on the Chandler,Nanushak,Oolamnagavik,and Killik Rivers,and old nests
on the Killik,East Fork of the Oolamnagavik,and in the Ivotuk Hills.
There are now enough records from the past five years to plot out the
northern limits of the breeding distribution on the Arctic Slope with fair
detail (Figure 2).The approximate latitudinal limit is 69°30'N.,although
some pairs can no doubt be found nesting somewhat farther north in the region
of the Sadlerochit Mountains.It is.significant,however,that no eagle has
ever been found nesting on the main Colville,although Cade did find an old
stick nest between the Killik and Umiat in 1956 that he thought lomB an eagle's
nest.Clearly the main breeding stronghold for this species in Arctic Alaska
lies in the Brooks Range and associated mountains,just as the Alaska Range
provides for a similar stronghold in the subarctic interior of Alaska.Non-
breeding in any given nesting territory is frequent (successful breeding may
occur on average only once in four years),and the montane distribution of
these nesting territories means that there is little overlap between breeding
eagles and breeding Peregrine Falcons.
4.The Rough-legged Hawk
RouSh-legged Hawks were nesting in near peak numbers in 1975,although
their productive output appeared to be only moderate.We observed a total of
162 pairs within the survey area,including one nest situated about 20 feet up
in a cottonwood tree with a crown of about 30 feet on the Chandler River.Along
the main Colville between the Etivuluk and Ocean Point,we found 68 pairs nesting
or attempting to nest;40 of these pairs were below Umiat.118ny of the nests
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were covered by incubating adults that refused to rise when the aircraft flew
by,so that we did not obtain much information on nest contents,but those that
we did see into contained mostly 2,3,or 4 eggs,only once as many as 5 eggs.
The average clutch size in 1975 apparently was not much more than 3 eggs.
l~en compared ~dth data for previous years on the main Colville popu-
lation,these figures indicate a return to peak numbers following a low in 1973:
In 1967,64 pairs laid an estimated 243 eggs,with a mean clutch of 3.95 eggs
(83 eggs observed for 21 pairs),and they fledged an estimated 122 young (69
young observed for 47 pairs),an average of 1.91 young per pair.In 1968,59
pairs laid an estimated 209 eggs,average of 2.19 per pair (92 eggs observed
for 42 pairs,including 13 with nothing),and they fledged 84 young,an average
of 1.42 per pair.In 1969,89 pairs (an all-time high)laid an estimated 303
eggs,average of 3.40 per clutch (235 eggs observed for 69 pairs),and they
fledged an estimated 160 young,an average of 1.80 per pair (79 young observed
for 44 pairs).In 1971,74 pairs laid an estimated 201 eggs,an average clutch
of 2.72 eggs (160 eggs observed for 54 pairs),and they fledged an estimated
102 young,an average of 1.39 per pair (96 young actually observed for 69 pairs).
In 1973,J.R.Haugh could find only 26 pairs of Rough-legs between the Etivluk
and Ocean Point,with none present between the Killik River and Umiat }fountain,
an all-time lm~for this population.Microtine rodents were at a low point in
their population fluctuations in the same year.Like the Gyrfalcon,the breed-
ing population of Rough-legged Hawks varies somewhat from year to year,and so
does its productive output;but the species continues to thrive over the long
term in the arctic and,indeed,is the most abundant falconiform during the
summer months.
5.The Raven
This passerine species is functionally a raptor and a cliff-nester,
so that it logically should be included in any consideration of cl1ff-nesting
birds of prey.Ravens also bred in maximum numbers along the Colville and its
tributaries in 1975.We found six nests on cliffs along the main Colville,and
one pair was also nesting in the rafters of the old SeaBee building at Umiat.
Eighteen other nests were found on tributaries and along the upper Colville
above the Etivluk.We were also told about a pair of tree-nesting Ravens that
used a large willow on the Ivashak River in 1967 (Harry Quinn,verbally).
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DISCUSSION AND RECOMMENDATIONS
A.THE USE OF FIXED-rITNG AND HELICOPTER AIRCRAFT
We have rather mixed feelings about the use of aircraft in work of this
sort.Clearly we are able to cover a much wider geographic area and to do so
in much less time.We are alAo ab19 to a~amfnp-Romo of the more remote cliffs
that might not othervnse be visited.We are unconvinced that we can be as
thorough with aircraft as we can be on the ground,especially in unfamiliar
terrain.Even in well-known areas thoroughness is hard to achieve at certain
times in the breeding cycle,especially just prior to ~gg llatching,when the
female is reluctant to flush even when approached on foot.This problem was
graphically illustrated this summer by comparison between our survey of the
Yukon River and the results obtained by Ambrose and Ritchie.Certainly one
factor that influenced our missing some pairs on the Yukon was the use of a
fixed-wing airplane.With a helicopter we might have achieved better results.
Generally,fixed-wing aircraft are excellent for survey work on Gyrfalcons,
Golden Eagles,and Rough-legged Hawks and may also be for Peregrines after the
young have recently fledged and can thus be flushed from the cliff,increasing
chances of observation.By that late in the year,however,an important segment
of the population--the pairs that have failed--are missed and are not accounted
for •
We have yet to see any behavior that convinces us that the prudent and
routine use of aircraft is harmful to breeding raptors.Any adverse effects of
low-flying aircraft,etc.,would be hard to demonstrate without long-term
controlled reproductive and population studies.About half of the Gyrfalcons
we approached at nests in the Helio Courier attacked without hesitation and
showed no fear or fleeing reaction.Others continued to feed their young without
looldng up as we flew by to check the eyrie.l-Iany Peregrines could not be flushed
from perches or nest sites by either fixed-wing or helicopter and often left
only after we were on the ground and out of the aircraft.Rough-legged Hawks
sitting on eggs almost never got.up when we flew by.
Our general recommendation would be to use aircraft for preliminary work
when an overview of the terrain,the general population dispersal and density,
etc.,need to be determined or when order of magnitude estimates are sufficient.
When precise data are necessary for population status and reproductive perfor-
mance,even in well-known areas,on-the:-ground ..work .is·.·.preferable.
To conclude,birds of prey are not importantly threatened or significantly
influenced by men flying around in airplanes or helicopters,as long ~they
remain in the air and do not deliberately harass the birds or shoot them.l~at
can threaten birds of prey and can have an adverse effect on their reproduction
and survivorship are the activities of men ~the ground.We feel that recent
state and federal regulations have placed too much emphasis on restricting the
use of flying aircraft and have not sufficiently considered needed restrictions
on the ground.These restrictions should consider both temporary and permanent
actions.The former category should include such things as the activities of
geological survey parties,geophysical teams,and any other scientific or inves-
tigative groups that have work to do in the environs of active nests.Generally
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the most sensitive times for disturbance are prior to egg-laying and during
incubation.Once the eggs have hatched,the birds can tolerate considerably
more activity around their nests without adverse effects.All sorts of develop-
ment and construction projects should be included in the latter category--
roadwork,bridge construction,pipelines,powerlines,installations,borrow
pita for gravel,etc.
It is important to recognize that certain specific cliffs,certain specific
trees,certain specific hillsides are nesting habitats for birds of prey.These
specific sites are used over and over--year after year--to the exclusion of
numerous other,seemingly identical places.This phenomenon of "site tenacity"
has been dramatically demonstrated by the Peregrines in Great Britain during
the build-up phase of the breeding population there since the low point reached
in 1963.The same old cliffs that had been occupied by pairs before the post-
lvorld Har II decline are the ones now being taken over by the new recruits to
the breeding population (Ratcliffe,1972).No new cliffs have been occupied.
Every time one of these traditional and historical nesting places is made
unsuitable for falcons by any of the limitless forms of environmental degra-
dation associated with man's activities,there is one less opportunity for a
pair to nest,and the species is consequently reduced by one breeding pair.
This is the reason why it is so important for the future of Peregrines and
Gyrfalcons--and all cliff-nesting species--to preserve the natural and biolog-
ical integrity of as many of these nesting cliffs as possible.
There are at least three specific places along the pipeline route where
this principle is being disregarded,although it is gratifying to note that the
Franklin Bluffs are one area where it has been applied for the welfare of nest-
ing Peregrines.
B.THE "OLD MAN"AND "SLOPE MOUNTAIN"SITES FOR FALCONS
We are concerned that earlier in studies of the pipeline more emphasis
was not placed on sensitive species (such as Gyrfalcons)in addition to endan-
gered species.Perhaps we could have influenced the choice of sites for the
borrow pits in the "Slope 110untain"area where Gyrfalcons nest.Another site
might have been selected to avoid working directly under the falcons.However,
we now recommend that the use of "Slope 110untain"by Rough-legged Hawks and
Gyrfalcons in the area of the borrow pit be carefully documented for the next
few years.This situation could yield important data on the tolerance level of
some raptors to disturbance by heavy equipment.Additionally,we learned in
October 1975 that Marvin Mangus had found a Peregrine nesting on the cliffs of
Slope Mountain in 1963,and it thus appears to have been an historical site for
Peregrines as well.
It is also unfortunate that the Gyrfalcons nesting on a small rock near
the road south of the Old Man Camp were not more regular in their occurrence,
so that we might have recognized the importance of that site earlier and influ-
enced the placement of the road.Depending on the amount of traffic,both air
and ground,that site may never be used again.W~know of an exact parallel in
Mount HcKinley National Park.For years Gyrfalcons nested on a small rock in
the Savage River Valley less than 1/4 mile from the park road.When a new
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bridge was constructed across the Savage River in 1951,the falcons deserted,
and the rock has never been used since.
We recommend that raptors as a group,especially falcons and eagles,be
recognized as sensitive species,rather than emphasizing only endangered ones,
when regulations relevant to the welfare of wildlife are contemplated for
development projects in Alaska.
Such recognition should provide a greater measure of care and protection
to those species likely to receive the greatest impact during this exploitation
phase in the Arctic.
C.THE "HOODOOS"
We also learned that there is a plan afoot between the State of Alaska and
Alyeska to construct a "tourist"road from the Alyeska Camp on the Richardson
Highway above Summit Lake to the foot of the Gulkana Glacier.This road would
run by the base of the "Hoodoosll--a rock formation that has been used.for
nesting by Gyrfalcons for at least 25 years and probably for much longer.
Ornithologists know this area as one of the best for Gyrfalcons in the State of
Alaska,and it is certainly one of the most scenic and spectacularly beautiful
falcon eyries anywhere in the world.Careful consideration needs to be given
to the impact of a road and tourist traffic passing'so close to this eyrie •
We recommend that any road stop at least one-balf mile away from the Hoodoos
and that tourists wishing to visit the glacier be required to·hike in the
remaining distance on a trail that keeps away from the Hoodoos proper and
directs foot traffic up the right limit of the glacial outwash.
D.THE "CRITICAL HABITAT tl CONCEPT AND RAPTORS
Allied to the recognition of sensitive species is the designation of
parcels of environment as habitat critical to the biology of those species.
We recognize that in Alaska cliffs that are now occupied or have been histori-
cally occupied by Peregrines and Gyrfalcons are critical habitat.We also
recognize that a buffer zone around those cliffs is important as critical
habitat.That zone probably varies with the sensitivity of the pair involved
and on particularities of the habitat in question.Perhaps it might vary
between 500 feet and one-quarter of a mile or in some cases exceed a mile.
What to do about the habitat necessary to support the pair with food is still
an open question.It is nonetheless critical habitat.We suggest that all
Peregrine and Gyrfalcon cliffs on federal and state lands be declared critical
habitat along with a buffer zone at least one-half mile in radius.
E.IHPORTAl-lCE OF THE COLVILLE AND YUKOli HATERSHEDS
It is clear to us that falcons occur in·maximum density in certain areas
only in Alaska.Other areas adjacent to them may have scattered populations,
but the major river is the "center"of the regional population.The Yukon and
Colville Rivers are two such rivers.So also are the Porcupine River and Tanana
River.We also recognize that some of the tributaries may have numerous pairs,
such as the Charley River possibly;but the main river valleys appear to provide
r---\the optimum habitat for these regional populations,and the pairs nesting along
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References
Bishop,L.B.1900..~otated list of birds.In W.H.Osgood,Results of a
biological reconnaissanc.eofthe Yukon ltlver region.Nortb Amer.Fauna
no.19.
Cade,T.J.1953.The peregrine falcon in the YultonV8lley.Proe.Second
Alaska Sci.Conf.1951:'330,·,33/,.--
...""a''',-....."."".~I,';ll :l,c···".,,~;d ',.~.~--,.!'~""!':'l"
Cade,T.J.1960.Ecology of peregrine and gyrfalcon populations in Alaska.
Univ.calif.Publ.Zool.63(3):151-290•.
Cade,T•.1.,J.L.Lincer,C.M.lo1hite,D.G.Roseneau,and L.G.Swartz.1911-
DDE residues and eggshell changes:inAlaskan falcons and hawks.Science
112:955-951.··.
Cade,T •.1.,C.M.lfllite,and J.R.Haugh.1968.Peregrines and pesticides
in Alaska.Condor 70(2):170-178.
Enderson,J.H.,D.G.·Roseneau,and L.G.Swartz.1968.Nesting performance
and pesticide residues in Alaskan and Yukon peregrines in 1961.~85:
683.
Enderson,J.H.,S.A.Temple,and L.G.Swartz.1912.T1me-lapsephotographie
records of nesting peregrine falcons~Living Birdll:113-128.
Fyfe,R.,S.A.Temple;and T.J.Cadet 1976.The 'North American Peregrine
Survey,1975.Can.Field Nat.In press.
Haugh,J.R.
Alaska.
1976.Recent history of peregrine falcons in the Tanana Valley,
(Manuscript submitted to Can.Field Nat.)
~
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Hickey,J •.1.,1942.Eastern population of the duck hawk.Auk 59:176-204.
Kessel,B.and T.J.Cadet 1958.Birds of the Colville River,northern
Alaska.Biological Papers of the Univ.of Alaska no.2.
Peakall,D.B.,T.J.Cade,C.M.l-lhite,arid J.R.Haugh.1915.Organochlorine
residues .in Alaskan-peregrinel3.Pesticides l.fonitoring Journal 8:255-260.
Ratcliffe,D.A._1912.The peregrine population of Great Britain in 1971.
Bird Study 19(3):117-'156.
Temple,S.A.,J.H.Enderson,and L.G.Swartz.1910.The upper Yukon Valley
of Alaska.Page 293 in The North American Peregrine Survey,1970,edited
by R.Fyfe and T.J.Cade.Can.Field Nat.84(3):231-246.--"White,C.H.1969.Breeding Alaskan and arctic migrant populations of the
peregrine.In:Peregrine Falcon Populations,Their Biology and Decline,
edited by J.J.Hickey.Univ.t-1isconsin Press,Madison..
White,C.1-1.and T.J.cadet 1971.Cliff-nesting raptors and ravens along
the Colville River in arctic ~M~Living Bird 10:107-150.
U.S.Fish &Wildlife Service
1011 E.Tudor Road
A~chora~~l Alask~995~