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Background Information
M Several basic assumptions made in the development of the original
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~Susitna Hydroelectric Anadromous Adult Scope of'Work may,in view of recent
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~8data analysis,be questionable.Two areas of major concern which may
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~result in misleading data are assumptions relative to the capabilities of
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Mthe side scan sonar and the application of mark~recapture techniques for·
SUSITNA RIVER HYDRO STUDIES REWRITE
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enumeration of salmon spawning populations.
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the original design of the adult salmon program was that comparative analysis
of fish abundance between sonar sites could be made.Onsite counts could
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-then be compared,and estimates of between &ite and maj~tributary spawning
populations made.This is technically incorrect for most Susitna stocks.
The complexity of the Susitna system (glacial,fluctuating water levels,
Illultiple channels,wide cross section,and be~l.~;:~~[n of salmon species)
render sonar counts as indices of run strength rather than absolute counts.
Data from three years of side scan sonar use (1978-1980)support the concept
that sonar counts are useable only as :un strength indices for chum,coho,
and king salmon.These same data indicate that count~attributed to sockeye
and pink salmon may be reasonabl,~estimations of spawning population sizes.
Figures 1 and 2 show the horizontal distribution for sockeyes and pinks
along the sonar substrate'in 1980.It is clear that both species tend to
use the bank in upstream movement in the Susitna River.Conversely,Figure 3
shows chum movement in 1979 to be more evenly distributed along the substrate.
This is an indication that chum counts are minimum numbers and that an
unknown component of the chum escapement is probably moving outside the
60 foot range of the side scan sonar.
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Fish targets attributed to coho salmon have not been plotted in a
similar manner because in both 1979 and 1980 there were not time periods in
which coho counts represented more than a small portion of the total salmon
run,and therefore coho horizontal distribution was masked by that of other
more numberous salmon species.However,other evidence exists which supports .~D~.I"W
the idea of offshore movement or fishwheel avoidance by coho sa1mo~The
attached memo (Appendix I)from Dave Watsjo1d (Sport Fish Division,Palmer)
indicates that if Commercial Fisheries Division escapement totals for coho
in 1979 are considered absolute counts,approximately 46%of the run was
harvested by sport fishermen above Susitna Station.As he pointed out,
this figure is unrealistic in view of the fact that only a small portion
of the Susitna river tributaries are accessible to fishermen and most of
in 1980 if sonar counts attributed to coho salmon are considered to be
times,indicating the same or greater percent of run harvested by fisherman
these are on the east side of the mainstem Susitna (data from 1979 and 1980
suggest that most of the coho may migrate up the Yentna River).Although
1980 sport fish harvest data are not yet completely tabulated)Sport Fish
lDivision personnel in Palmer feel that the harvest may exceed 1979 by 2-3
In 1980,the harvest
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Kenai River coho catch and escapement totals demonstrate f~
'\''"\6TIt)\')l,i,;1/the error in assuming coho counts are absolute.
absolute counts.
exceeded 6,000 while fish targets attributed to coho at the sonar site were
less than 8,000.
In 1978 a special appropriation was made available to the Oivision of
Commercial Fisheries to use side scan sonar to enumerate king salmon in the
Susitna River.The data indicate offshore movement by larger and older
king salmon.Fish targets attributed to king salmon numbered approximately
43.000 at Susitna Station while S~ort Fish Division aerial surveys'produced
an estimated escapement of 74,000 for the Susitna drainage.Age information
.....
from 1978 also indicate that larger kings move in mid-river.Susitna
Station fishwheel catches yielded 48%four year old and 30%five year old
kings.Sport Fish Division data from the Deshka River (where roughly half
of the total Susitna River kings were counted)shows that the two most
numerous age classes were five and six year old with 42%and 38%of the
fish sampled.respectively.
The above data on chum,coho.and king salmon clearlY shows that the
assumption that counts for specific mainstem and tributary salmon ·spawning
populations can be arrived at by subtraction of counts between the sonar
sites is invalid.Counts at each sonar site will represent only the minimum
number of these species which pass the site.and will not represent total
population counts,or indicate the extent of mainstem spawning.In the
case of sockeye and pink salmon •.counts may be close representations of the
actual spawning populations.but still do not indicate the degree of
mainstem spawning.
The existing Susitna River Hydroelectric Project scope of work also
outlines the use of a mark-recapture program to enumerate salmon above the
Sunshine sonar site.The basis of the program is that tag recoveries will
give salmon population estimates for the Susitna River above Talkeetna (to
Devil Canyon),which can then be subtracted from Sunshine sonar counts to
get the numbers of salmon going up the Chulitna and Talkeetna Rivers.This
program,however,is based on several questionable assumptions about mark-
recapture programs.rna i nstem spavmi n9,and the abil ity to use sonar counts
as absolute estimates.
There are seven assumptions which must hold true for accurate population
estimates using any mark-recapture technique.These are listed below,
accompanied by data from the literature detailing how assumptions were.
~tested in other salmon mark-recapture programs,and the effect on population
.....
"estimates in the event that assumptions are not met.Existing data from
Susitna River studies are also included where possible to document problems
which could occur in meeeing the stated assumptions.
1.Loss by natural mortality or emigration i$proporttOna11y the
same for marked and unmarked fish.
The net effect of a lost of marked fish relative to unmarked
fish is an overestimation of the total population.Schaefer (1951)
documents previous work which found that gill nets are biased
toward externally tagged fish.He also.suggests (and cites other
authors)that fish already stressed (in his worK this was indicated
by gill net marks)are more susceptable to mortality after handling
in the tagging process.Although difficult to assess,the stress
affect on sa1~on could be significant in the Susitna River due to
the high percentage of fish with recorded damge by nets,predators,
and parasites.Data from 1974,1975,and 1977 (AbF&G,1977)base-
line Susitna River work indicate that loss by emigration could be
a significant factor in developing population estimates for that
portion of the river above the Talkeetna River-Chulitna River
confluence.These studies found that a portion of the fish tagged
on the Susitna River above Talkeetna were later recovered in other
downstream tributaries.Conclusions in these reports suggested
the confluence may be a milling area,however the possibility
exists that the 'drop out'phenomena could be a reaction to the
tagging process.In either case,the consistant appearance of
numerous tagged fish below the tagging sites is an indication
that population estimates may be biased if the behavior is not
true for the unmarkedpopulatfon of salmon.Schaefer also con-
eluded that even if the ratio of marked fish to total fish is
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"...
2.
assumed to be the same at the mark and recapture sites,the
estimate of the size of the population is correct only for the time
(and place if migratfon occurs)of marking.This.is true especi-
ally for situations where marks are put on salmon at a distance
from the recapture area and an unknown rate of mortality (or
migration up tributaries or into side sloughs)equal on both marked
and unmarked fish reduces the population enroute to the area where
recapture sampling is done.In such a case the population estimate
is correct for the locality of marking only.In the Susitna River
this would mean that a mark-recapture program where fish were
tagged at the Sunshine site would yield an estimate of the siz~of-
the run which passed the Sunshine site only (where sonar counts.
are also being conducted)~
Marks are not lost during the experiment.
None of the literature reviewed or previous Susitna work
indicates a problem with tag loss.
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3.Marks are recognized and reported on recovery.
This is not a potential problem because recaptures in the
ttt-upper Sjsitna River will be made by project personnel.The project
does not rely on tag return from the sport or commercial fishery,
a source of error in many studies.
4.Marked fish are as likely to be caught as unmarked fish.
In order for population estimates to be valid using standard
statistical approaches,the ratio of marked fish to the entire
population must remain constant during recapture.This assumes
that the recovery method does not discriminate against either
marked or unmarked fish.Ricker (1978)documents known cases
where fish behavior and cathability can be altered by fish
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handling,which may reduce the number of recaptures of tagged
fish.Conversely the high visibility of tags on live fish may
increase their chances of being counted in water conditions where
"\unmarked fish are marginally visible.In any event,Schaefer
(1951)demonstrated that the use of periodic spawning ground
surveys for counting tagged and untagged fish was not sufficient
to assume equal catchabi1ity of the marked and unma!'ked fish
when sites were long distances apart.Work compiled and ed,ted
by Ulland (1977)found that the assumption is most valid (error
reduced)when relatively large numbers of recaptures can be made
in a relatively short period of time.
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5.Marked fish mix randomly with unmarked fish,or fishing effort is
proportional to the density of fish in different parts of the body
of water.
Although random mixing of marked and unmarked fish is difficult
to assess,Schaefer (1951)found that there was not full mixing of
sockeye even when all fish were equally susceptible to capture and
recovery.As noted previously,evidence exists from earlier
Susitna River work that kings,chums,and cohos are undercounted
because of offshore movement or fishwhee1 avoidance.Without any
means of assessing the magnitude of the runs of these,species
which are not captured by fishwheels it may be impossible to
determine the degree of mixing,or to assume that fishing effort
is proportional to fish density in the river.
Recapture methods concentrate on four major tributaries and
the mainstem sloughs in between.It can be assumedfrorn the
literature that mi~ing has not been totally random,however no
means exists to assess the degree of error which may result in
--------~---------~----------~-
population estimates.Limiting of recapture sampling to side
sloughs and tributaries within a relatively small section of the
upper Susitna River leaves researchers without any means of
determining if effort is proportional to fish densities in differ-
ent parts of the river.In fact,a major portion of the river
and within the river has been left out of the recapture sampling
effort indicating that unless the unsampled portions of the river
contain no fish,population estimates will be in error.
6.A negligtble amount of recruitment occurs during the period
between marking and recapture.
Potential sources of error in population estimates exist fn
the current study plan because there is no way to.assess recruit-
ment in the recapture areas.Fish which move outside the fishwheel$-I
in the lower river may move into sloughs and tributaries where.
they are countable in recovery sampling only.As stated previously
emigration may occur after tagging,giving a form of rev~rse
recruitment which could affect estimates.Lastly,limitlng the
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area of recapture sampling offers numer9us combinations of tagged
and untagged fish movements and spawning area selections which
could result in recruitment in recovery sampling.'Because of the
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limitations of the existing program.and the complexity and enormity
,
of the spawning area in the upper Susitna it will be extremely
difficult if not impossible to assess the affect of recruitment on
7.
population estimates.
The product of the number of fish originally captured and marked,
and the number of fish in the recapture sample examined for marks
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must exceed the product of the size of the popuration at the time
of release of marked fish times four.
Although the number of tagged fish recaptured can be low (if
the number of fish originally captured a~d marked divided by the size
of the population at the time of release of marked fish is small,
statistical bias can be ignored if the number of recaptures exceeds
four)a fairly large number of fish must be tagged at the lower
site and captured at the upper site to give valid estimates.Fo~
example:if 100,000 pinks are suspected to enter the upper Susitna
River above Talkeetna,then 500 fish would need to be marked and
•200 examined for marks.Any combination of multipliers of marked
fish and fish examined for marks which yields a product greater
than 100,000 would be a sufficient sample size.If,however,
precise estimates of spawning populations are required,much larger
sample sizes will be needed.
Robson and Regier (1964)developed charts which give the minimum
sample sizes for marked fish and total recaptures (marked plus un-
marked)necessary when an estimate of the total population is known.
Using these charts and preliminary estimates of the run sizes into
~the Susitna River above tIe Yentna River confluence it is possible
to develop sample sizes which would be needed for a statistically
unbiased population estimate (with a 95 percent confidence that
errors will not exceed 10 percent).
Case 1.All Fish passing the Susitna-Yentna confluence will also
pass the Sunshine Site.
Sped es Year
Estimate of
Population Size
Number
Marked
Number Examined
For Marks
Using these data and Susitna River fishwhee1 capture totals -
Pink 1982
Sockeye.1982
Pink 1981
5,000
6,000
3,000 f.l'
4,000
,,1 06,000 qJ .
6,000
7,000
12,000
2,000
2,000
/
4,000
5,000
11 ,000
4,000 /
'I~2,000
50,000
70,000
20,000
100,000 /'
1,500,000
1981
1982
1981
1982
1981
Coho
Coho
Chum
Chum
Sockeye
...,.
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from 1978-1980 it appears that as many as 10 fishwhee1s would be
necessary to tag enough fish.If only 10%of the estimated 1981
and 1982 runs were to reach the.Sushine Site it would still be
necessary to run three or four wheels to tag the minimum number of
chum salmon (the species thought to be least catchable by fish-
whee 1s).
The problems discussed above indicate that the mark-recapture
program as it now exists has the potential to give misleading
information about the spawning salmon populations above the Sunshine
sonar site.The numbers generated could easily under or over esti-
mate the upper Susitna salmon escapement (Talkeetna to Devil Canyon)
which when subtracted from Sunsmne Sonar counts,w~ich are assumed
to be abs91Yi~,will also give misleading results about the sus-
"",-"_.-,_m<_.~'__-·~·
pected escapements into the Chulitna and Talkeetna Rivers.
With the existing program problems in mind,several different
approaches to obtaining appropriate data about each salmon species
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are discussed below.As background some basic information which is
known or suspected about the run tendencies of each species in the
,
Susitna River are pointed out,the questions which need to be
addressed by the program are 1i sted,and programs-'are presented by
species which are felt to be the best means of answering the
questions within the time,economic,and technical limitations
that currently exist .
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•t
'REVISED PROGRAM
The following section presents a revised anadromous adult program
designed to meet,where possible,the objectives stated in the existing scope
of work.Each salmon species is freated individually in terms of sampling
methods and the degree of success which might be achieved with the revised
program.Selection of methods and evaluation of program success incorporate
run expectation data and professional judgements expressed in Appendix II.
OBJECTIVES
The objectives stated in the existing scope of work are br6~enddwn
into three main study areas,and are the same for all species within each
area.All of the objectives.howeve~.can be condensed into five basic
objectives which apply to all species as follows:
1.Escapement data by species in total and for each study area of
the river from Susitna Station to Devil Canyon;
2.Differentiation of the Susitna River and Yentna River contributions
to the total escapement;
3.Run timing into each study area of the Susitna River;
4.The degree of mainstem spawning in the designated study areas of
the river,and
5.Differentiation of Chulitna,Susitna,and Talkeetna River stocks.
Individual programs are discussed for each objective by species in the
following section.
Sockeye
1.Side scan sonar counters will be placed in the Susitna River at Susitna
Station,the Yentna River,the Susitna River at Sunshine,and in the
Susitna River above Talkeetna.The counters should provide acceptable
estimates of sockeye escapement at the respective sites.It is likely
.,..,
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that sockeye (and pink)will be the only species for which the
'subtraction'method of obtaining counts between sonar counters is
va 1i d.
A fin and tail clipping program will be used at Susitna Station to
determine if east bank sockeye go up the Yentna River,or west bank
sockeye move up the Susitna River past the confluence.Clipped fish
recoveries which indicate cross over will not provide absolute counts
of the number of fis~,but may indicate the degree in a broad sense
(i.e.minimal versus significant).If cross over does not occur it can
be assumed that Susitna Station west bank sockeye counts are fish migra-~15
,,-.:t1fJ1 ~ti-l ~Il·./
ting up the Yentna River.Yentna sonar sockeye counts can then be
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compared "to Susitna Station west bank co~ts to deter~~t~e.!._
'subtraction'method is feasible --t<J,;)~-wU.1~~~•«-#-~c.,..~~.
3.Run timing data will be compiled from fishwheel captures at all sites.
4.Aerial surveys will be conducted of the known clear water sockeye
spawning areas of the Susitna River above the confluence with the
Yentna River ..Surveys will be conducted a minimum of three times to
determine peak abundance of sockeye.Aerial counts can be compared to
sonar counts to give an estimate of the maximum number of potential
mainstem spawners.It should be stressed,however,that surveys of this
type usually do not account for one hundred percent of the spawni n9
population.Previous surveys from the Susitna Basin have enumerated
from 10%to 30%of the estimated sockeye escapements.Increased survey
time,increased number of survey areas,and complete surveys of existing
spawning grounds (fixed boundaries exist in each established survey
area whi ch 1 imit the area of survey for year to year consi st;-~y)will
undoubtedly increase the number of tributary spawners accounted for.
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However,because of the enormity of the area and number of tributaries
in which small sockeye runs exist,it is doubtful t~at any amount of
effort will successfully enumerate all sockeye spawning in tributaries.
5.In the case of sockeye (and pink)salmon,placement of sonar above and
below the confluence of the Susitna,Talkeetna,and Chulitna Rivers
should give some idei!..,of the magnitude of the Talkeetna-Rtvi:1"-Chulitna-
_River run.Aerial surveys may help determine how this component breaks
down within the two rivers.
pinks
1.See sotkeye number one .
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4.
See sockeye number two.
See sockeye number three.
A survey of historical data will be undertaken to determine 'if an aerial
program such as the one presently proposed for sockeye can be accomplished.
As with sockeye,spawning ground survey totals can be compared to sonar
counts to estimate the ma~imum number of mainstem spawners.In addition,
two ground survey crews will be placed on the mainstem Susitna River for
the purpose of identifying spawning areas in mainstem channels,sloughs,
'-----~bac~s and other areas where pinks have been obs"erved spawning in'/~_1..~__
_past years.With experience these crews may be able to give reliable ~_
estimates on the number of spawners in each mainstem area examined which ~/
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when totaljed will represent a rough estimate of the mainstem spawning
population which might be impacted by the proposed Hydroelectric project.
5.See sockeye number five.Ground survey crews will not be working the
/\
Talkeetna and Chulitna Rviers where it ;s suspected that some pink
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spawning will occur.
l).'.'
of the escapement will go up the Susitna River pas~the Yentna River;
and 4)inability to predict the effects of the fishery on the returning
population may increase or decrease the escapement.Based on these,..,.
assumptions and recognizing that any estimates of return will be very
rough,the sockeye run into the Susitna River above the Yentna River
may approach 70,000 in 1981 and 65,000 in 1982.
Pinks
11.If the same assumptions about bank movement related t?direction of travel
at the Susitna River-Yentna River~onfluence are true,an average of
69.5%of the pink run will continue up the Susitna River (68%in 1979
and 71%in 1980).
2.Aerial surveys enumerate pinks incldentally to sockeye and as such,
~tributaries known for large pink spawning populations are not necessarily
surveyed.In 1980 pinks were countea in large numbers in tributaries
entering the Susitna above the Yentna in Chunilna Creek tributaries only
(the Chinulna enters the Talkeetna River).Visual observations by sonarr
crews in 1980 found large numbers of pinks spawning in every slough or
backwater investigated in the vicinity of Susitna Station.It is felt
that at least in even year escapements,a large part of the run spawns
in or near the mainstem river.
3"Surveys in 1977 enumerated 6,600 pinks (of the 1.5 million estimated
escapement)in the area above the three river confluence (Talkeetna
River,Chulitna River and Susitna River)to Devil Canyon.In 1975,291
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of the estimated 546,000 escapement were enumerated in the same area .
4.The 50%point of the pink run past Susitna Station historically oc~urs
around the end of the third week of July in odd years and the end of
July in even years.Data from upper Susitna Studies conducted in 1974
1.
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and 1975 indicates a peak in pink catches in the first two weeks of
August.
5.Predictions are based on these assumptions:1)the number of fish
returning to the river per parent year spawner was 1:1 in even years
and a range of .08-2.7:1 in odd years based on escapements at Susitna
Station from 1978 to 1980;2)70%of the total pink escapement continues
up the Susitna River past the Yentna;and 3)inability to predict the
effect of the fishery on the run may influence escapements.Based on
these assumptions and rec~gnizing that run predictions are very rough,
the Susitna crews above Susitna Station "may be dealing with pink run
sizes in the Susitna above the Yentna of 70,000 to 236,000 in 1981 and
1.5 million in 1982.
Chum
If the assumption that bank selection indicates direction of travel at
the Susitna Rivers confluence,82.5%of the chums are moving up the
Susitnaabove the Yentna.It should be noted,however,that because
of potential midriver movement by this species,the percentage attributed'
to each river may not be close to actual escapements.That is,the
figure represents the percentage of chums catchable by a fishwheel only.
2.Spawning ground surveys rarely account for more than a few chums in any
clearwater area,although occasional reports of large numbers exist in
the historical records (in 1953 10,000 chums were spotted off the mouth
-of Chunilna Creek,a tributary of the Talkeetna River).Susitna Hydro
surveys accounted for approximately 3,100 chums in 1974 and 2,300 in
1977 (of a total escapement estimated at 105,000 by Susitna Station)in
the Susitna River above the confluence at Talkeetna.It is suspected,-.
that chums can and do spawn in the bottom conditions of the mainstem
river.Most of the chums observed in the 1974,1975,and 1977 Susitna
Hydro studies were found spawning in side sloughs of the upper Susitna.
3.The 50%point of chum fishwheel catches at Susitna Station usually
occurs around the last week of July although the date fluctuates from
year to year and may be as late as mid-August (August 14 in 1979).
Chum captures in upper Susitna fishwheels peak during the period of
August 11-17 in 1974,1975,and 1977 studies.
~~.Predictions of 1981 and 1982 chum escapements are based on the'fol1owing
assumptions:1)totals are based on recent sonar counts which are thought
to be minimum escapement totals,2)the return is primarily 4 year old
fish;3)the number of fish returning to the river per parent year
spawner has averaged .6 for 1975 and 1976 parent years returning in 1979
and 1980;4)82.5%of the returning chum go up the Susitna River above
the Yentna;and 5)inability to accuratelY enumerate the run may result
in extremely low estimates of run strength (this is probably true for
1979 and 1980 escapement estimates,which also affects the return per
spawner ratio).Based on these assumptions and recognizing that
confidence in the above numbers is much less than for sockeye and pinks
the minimum chum escapement into the Susitna above the Yentna may
approach 50,000 in 1981 and 70,000 in 1982.
Coho
1.If the same assumptions above bank movement related to direction of
i travel at the Susitna River-Yentna River confluence are true for coho
as sockeye,roughly one-thi rd of the coho enteri ng the Susi tna River wi 11
continui above the confluence with the Yentna River.As with chums,
however,lack of reliable sonar counts greatly decreases the degree of
confidence in this figure.
\.._--"""----------,--~~---_._---_.~-~-
·2.Sport Fish Biologist feel based on experience that mainstem spawning
by coho salmon is negligible.No hard evidence to support this exists.
Aerial surveys rarely account for more than a few coho in clear water
tributareis,coho are the most difficult salmon species to enumerate on
spawning ground.The literature suggest that coho tend to spawn in
smaller clearwater creeks and rivers and are not known to use substrates
such as those typical of the mainstem Susitna River and its larger
tributaries.-I
3.Almost nothing is known about coho distribution either above the Talkeetna
River-Chulitna River confluence or in the mainstem between Talkeetna and
the mouth of the Susitna.
4.A 50%date for fishwee1 captures of August 15 in the Susitna River
above Talkeetna was established by the 1974 and 1975 studies,however,
this figure is based on a very small sample size.Fifty percent of the
cohos were captured in fishwheels at Susitna Station around August 1 in'
1979 and 1980.As with chum,these dates have varied by as much as two
F"
~weeks in previous years.
I
5.Predictions of coho run strength in 1981 and 1982 are very,rough because
the run strengths from previous years are probably low,giving a low
ratio of the number of fish returning to the river per parent year
spawner.The assumptions are:1)the run is primarily four year old
fish;2)the return per spawner ratio used is 1:1 although the yearly
ratio for 1979 and 1980 differs by nearly 100%and 3)the ?usitna River
contribution above the Yentna River confluence averaged 33%although as
explained above this figure represents a comparison of fishwhee1 captures
only and may not reflect the actual contribution of each river system.If
these assumptions are close,then the number of fish sampleable by current
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techniques and entering the Susitna River above the Yentna may approach
20,000 in 1981 ,and 30,000 in 1982.
King
Sport Fish input?
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APPENDIX II
RUN EXPECTATIONS
Based on previous experience on the Susitna River and analysis of 1978
through 1980 Susitna Station sonar and fishwheel data,certain characteris-,.
tics of each salmon species run can be described which have been used in
program redesign.Using these previous data and the assumptions contained
in the text for each species below,it 1s possible to give some information
about what answers might be expected and what information in unknown about
the questions posed in the scope of work objectives.
Sockeye
11.Because of the location of the sonar counters at Susitna Station just
below and across from the Yentna River-Susitna River confluence,it is
thought that most salmon following the west bank are going up the Yentna
River and those on the east bank continuing up the Susitna River.Some
unpublished data exist in support of this speculation in the form of
informal work done in 1978,as well as the 1977 tagging studies.No
fish captured and marked in east bank wheels in 1977 were later captured
in wheels located on the Yentna or Skwentna Rivers.In 1978,field
crews tail-punched sockeye in east bank wheels to determine if crossover
to the west bank was occurring,but of the fish marked only one was
later found in west bank wheels.The evidence is far.from conclusive,
(
but indicates that bank choice may determine the direction of travel
at the Yentna River-Susitna River confluence.Although no work of this
nature has been attempted for other salmon species it is assumed for
purposes of discussing relative magnitude of run up each river (Susitna
and Yentna),that the same pattern hold true for pink,chum and coho.
In 1979 and 1980 west bank sonar counts contributed 79%and 82%of the
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sockeye total respectively.The assumption that these fish went up the
Yentna River is somewhat supported by aerial surveys which enumerated
91%(1979)and 93%(1980)of all Susitna drainage sockeye spawners seen
in surveys in Yentna River tributaries.
2.Aerial surveys accounted for 27%of the west bank sockeye counts and
10%of the east bank sockeye in 15 known clear water tributary spawning
areas in 1980.These numbers may have been higher except for economic
constraints on survey time (particularly on the Susitna River above the
Yentna confluence),and 1.imiting of counts to established index areas.
Nothing is khownof sockeye mainstem spawning in the Susitna drainage
although it is thought that bottom conditions (primarily shifting sand
and silt)are not condusive to or preferred by sockeye for spawning~-
3.Baseline studies in 1974 and 1975 counted very few sockeye in the Susitna
River above Talkeetna.In 1974,336 sockeye were seen in surveys,and
in 1975 when 108,000 sockeye were estimated to have passed Susitna
Station,103 were counted in the upper river.Of the 238,000 sockeye
estimated at Susitna Station in 1977,ground surveys enumerated 661 in
the Susitna River above Talkeetna.
4.The date on which 50%of the sockeye run passed Susitna Station in 1978-
1980 was July 18-20.Fishwhee1s on the-upper Susitna River above
Talkeetna captured 50%of the total sockeye catch on August 5,1975
and 75%by August 16,in 1974.
5.Predicting returning spawning populations in difficult at best,but
numbers are generated here based on the following assumptions:1)The
return will be approximately one-half 4 year and 5 year old adults;
2)the number of fish returning to the river per parent year spawner is
2 to 1 based on 1980 returns from 1975 and 1976 parent years;3)20%
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Chum
1.Chum salmon (and coho)will be the hardest species to obtain accurate
escapement counts for.As~discussed previously,sonar counters are not .
jj~usable-.for absolute counts,and since most of this species are suspectey~//
.to be mainstem spawners,spawning ground surveys of clearwater streams
will probably not enumerate -a substa'ntia1 number.It is possible that
the majority of the Susitna River chum run continues up the mainstem
above the Yentna River to spawn,but very little data exists that docu-
ments where or how many_Ground survey crews will be used to document
those areas along the entire river where chums are fo~nd,and
various sampling methods will be used to obtain rough estimates of the
numbers of spawners in each area.This program is viewed as a preliminary
approach which may reveal areas where a significant amount of spawning
occurs that may be worthy of a concentrated enumeration effort in future
years.It is doubtful if the technology exists to enumerate chums which
spawn primarily in the glacial mainstem areas.This approach,however,
coupled with habitat flow work may roughly estimate the number of spawners
which may no longer have access to side sloughs when flow levels are
reduced.
2.Because of the difficulty in enumerating chums,differentiation of
Susitna River and Yentna River stocks will be accomplished only in the
form of minimum counts at the Yentna and Sunshine sonar sites.Although
east bank Susitna Station sonar counts attributed to chum are thought to
be migrating up the Su?itna River past the Yentna River confluence,
without accurate counts at any point in either river it will be impossible
to enumerate the percent of the total chum escapement contributed by
either river.
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3.Run timing data will be compiled from fishwheel captures at all sites.
4.See chum number one.
5.See chum number two ..Minimum chum escapement totals will be obtained
for the Susitna River above Talkeetna using sonar counts and ground
surveys.No estimates of chums migrating into the Chulitna and Talkeetna
Rivers will be made.
Coho
1.Sonar counters will provide a minimum estimate of co'ho passing each site.
Ground survey crews will report numbers of coho observed or captured
however since coho are not normally found spawning in the conditions
which exist in the mainstem Susitna River it is expected that ground
crews will not locate large n~mbers of coho.Apparently,coho ~re found
in nearly every clear body of water,regardless of size.in the Susftna
drainage,which makes large scale tributary survey efforts of questionable
value.
2.See chum number two.
3.Run timing data will be compiled from fishwheel captures at all sites.
It is felt that mainstem coho spawning is neg1igable;however,'no data.-
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4.
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exists to support this contention.
coho found in mainstem areas.
See chum number five.
VJ
Ground survey crews will enum~rate ~y
~:ing
1.Intensive aerial surveys will be conducted on known king salmon spawning
tributaries.This is an ongoing program developed by Sport Fish Division
to estimate total king salmon return to the Susitna Basin.
2.See king number one.
3.See king number one.
4.Aerial surveys are conducted on clear water tributaries only.No
attempt will be made to estimate the degree of mainstem spawning of
kings (if any)due to the glacial nature of the mainstem Susitna River
and it's main tributaries.
See king numbers one and four.
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L~._~__7 _
I •.1l.
Alaska Department of Fish and Game.1978 Preliminary Environmental
Assessment of Hydroelectric Development on the Susitna River.116 pp.
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Barrett,B.M.1974.An Assessment of the Andromous Fish Populations in
the Upper Susitna River Watershed Between Devil Canyon and the
Chulitna River.Alaska Department of Fish and Game Cook Inlet Data
Report No.74-Z.56 pp.
Friese,N.V.1975.Preauthorization Assessment of Anadromous Fish
Populations of the Upper Susitna River Watershed in the vincinity of
the Proposed Devil Canyon Hydroelectric Project.Alaska Department of
Fish and Game,Division of Commercial Fisheries,Anchorage,Alaska.
:120 pp.
Gulland,J.A.ed.1977.Fish Population Dynamics.John Wiley and Sons.
New York.372 pp.
Ricker,W.E.1975.Computation and Interpretation of Biological Statistics
of Fish Populations.Bulletin of the Fisheries Research Board of
Canada.Number 191.382 pp.
Robson,D.S.and H.A.Regier.1964.Sample Size in Petersen-Mark-
Recapture Experiments.Transactions American Fisheries Society.
July 1964.Vol.93,No.3.215-226.
Schaefer,M.B.1951.A Study of the Spawning Populations of Sockeye Salmon
in the Harrison RfverSystem,with Special Reference to the Problem of
Enumeration by Means of Marked Members.International p,acific Salmon
Fisheries Commission.Bull.4:207 pp.
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~AE~v10RANDUM State of Alaska
TO:
Bruce King
Commercial Fish Division
ADF&G.Soldotna
FROM:<7uJ·Davi'd'~~atsj old
Sport Fish Division
ADF&G.Palmer·
DATE:
FILE NO:
TEL:EPHONE NO:
SUBJECT:
November 4.1980
745-3178
Coho Data
Your memo was referred to me since I do all the anadromous fish work in the valley.
We do not monitor coho escapements "in very many streams due to the inaccessibility
and large numbers of coho systems that exist in Upper Cook Inlet.We do attempt to
maxe escapement counts on a small number of streams and we only count certain sec-
tions of these streams.Unfortunately the last two years have been extremely wet
~and complete counts have not been possible on some of the streams.The following
are those streams pn which we have some coho data:
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1975 1976 1977 1978 1979 1980
Wasilla Creek 207 313 *150 248 *
Cottonwood Creek 236 204 117 264.*870
Birch Creek 92 27 96 103 120 121
Question Creek 111 126 87 45 384 321
Fish Creek (Weir)1.601 765 .930 3.121 2,511 8,832
*Allor part of index areas not countabl e.
Birch and Question Creeks are the only two streams that are tributaries of the
Susitna River while the remaining streams flow directly into Cook Inlet.
Fish Creek and Cottonwood Creek are the largest coho systems that are monitored on
a regular basis.The 1980 escapements on these streams were the highest recorded
since Statehood.We do not census anglers on any of the east side·Susitna River
tributaries.but we are in close contact with the sport fisheries that occur on all
east side tributaries and streams flowing directly into Knik Arm.I can safely say
that in my eleven years of close association with the sport fisheries in this area
tha.t the excellent coho salmon fishing experienced in 1980 was unequalled in the
previous ten years.The apparently large number of coho available in 1980 was not
restricted to just several areas.but was widespread throughout all the Upper Cook
Inl et drainages..
Based on the 1979 sport fish catch estimates for the Susitna River,Larry Engel and
I believe that the estimate of 37,000 is well below the actual escapement level of
the Susitna River.We have received the preliminary sport fish·catch estimates from
the 1979 Statewide harvest study.In 1979 the coho catch estimate forSusitna River
tributaries was 17,000.If the Susi tna River coho escapement was indeed in the
range of'37 ~000 that would mean the sport fishery would have harvested 46%of the
escapement.This is not very likely in this system where the majority of coho strearr
are not even fished because of their remoteness.Even in the most ideal .situations
on our roadside stream systems we seldom see harvest levels that reach this magnitude
We also feel that the 1980 escapement.level of 43 ..000 is also a very conservative
fi~lre.ihe 1980 escapement in the Susitna River drainage could easily be twice the
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Coho Data -2-November 4~1980
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1979 figure given the fact that in Cottonwood and Fish Creeks the 1980 escapement
was 3.S times the 1979 escapement level.The sport fisheries also indicate a con-
sider~Jly greater abundance of coho in 1980 than what occurred in 1979.When the
1980 Statewide harvest study is completed sometime next summer I am confident that
it wilJl show a very significant increase in the coho harvest in all areas of Upper
Cook Inlet.
We have just bits and pieces when it comes to length-weight'averages and age
classes.Several years ago we collected length-weight data from Wasilla Creek,
Fish Creek and Little Susitna River.Wasilla and Fish Creeks coho are representa-
tive of the size coho that are generally available in Upper Cook Inlet.These
coho average 58-60 centimeters fork length or 53-55 centimeters mideye to fork and
4.6 to 4.9 lbs.The Little Susitna River coho have always been unique in that they
are larger than normal coho for this area.They average 67 centimeters (fork
length),60 centimeters (mideye to fork)and 7.9 Ibs..
The majority of coho salmon that have been aged from various systems in the area
have been Age 2.1.
I hope this information assists you in some manner.Unfortunately we donlt have
much information on coho at this time.If you have any questions regarding any-
thing that I have stated please contact me.