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PRE-AUTHORIZATION ASSESSMENT
OF THE
SUSITNA RIVER HYDROELECTRIC PROJECTS:
A LIMITED WILDLIFE STUDY
GAME DIVISION SECTION
BY
CARL MCILROY
GM1E BIOLOGIST
AND
TED SPRAKER
GAME BIOLOGIST
ALASKA DEPARTMENT OF FISH AND GAME
GAME DIVISION
ANCHORAGE
INTROillCfiON
PROCEilJRES
RESill.TS
TABLE OF CONffi\ffS
.lvloose Distribution, June 1974
Moose Wintering Distribution, 1974-75
Evaluation of Moose Wintering Range,
Moose Conditions , Loss of Winter
Range by Inundation.
Caribou Distribution and Trails
Harvests and I Iunting Pressure
Observation of other Marmnals
DISCUSSION A~ CONCLUSIONS
MITIGATIVE ACTIONS
REFERENCES CITED
Page No.
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UPPER SUSITNA RIVER \HLDLIFE STUDIES
by: Carl Mcilroy
Game Biologist III
Alaska Department of Fish and Game
INTRODUCTION
Reconsideration of portions of the Susitna River as a source of
hydroelectric power has necessitated a reevaluation of the effects of a
dam or dams on the area's indigenous and transient wildlife. Former
studies included an evaluation of the monetary values of the Susitna
basin based strictly on estimated harvests (Anon. 1954). However, the
applicability of those data to the present is limited because of changing
harvest patterns and changing calculations placed on an animal's worth.
A detailed report on the fish and >vildlife resources of the Susitna
basin and the impacts of the proposed Devil Canyon and Denali dams on
those resources (Anon. 1960) was an excellent evaluation considering
the limited information available at that time. This report is intended
to supplement the 1960 study by updating inventory and harvest data, by
reporting on big game distributions observed during the spring of 1974
and the winter of 1974-75, by reevaluating the main effects on wildlife
caused by the proposed Devil Canyon and \vatana Dams, and by suggesting
mitigating actions and future studies based on the current perspective.
PROCEDURES
Hoose distribution surveys during June, 1974 were flown with a PA-
18 supercub by ADF&G biologists. The Susitna River above the proposed
Devil Canyon Dam up to the Susitna Glaciers and the lower portions of
its major tributaries were surveyed (Fig. 1). Flight lines within the
surveyed area were approximately one mile apart, representing a survey
of moderate intensity. Big game distributions during the winter of
1974-75 were assessed by making five aerial surveys over the Susitna
study area at roughly monthly intervals. The Susitna study area for
these flights was defined as the Susitna River upstream from Gold Creek
and the lower portions of the Susitna River's major tributaries (Fig. 2),
Observations of all larger mammals '"'ere recorded, and those observation
numbers were located on a map. The upper limit of surveys was the
highest elevation that moose were found. The initial flight during
November was intensive, and moose sex and age composition were obtained
along with big game distributions. Complete subareas were searched for
moose. Because of poor weather, decreasing daylight, and increasing
ratios of ferry time to count time, not all of the study area was surveyed.
Subsequent flights, from January through April (Fig. 3-6), were less
intensive, and roughly fixed flight patterns were flown with no
attempt to search all subareas for moose. The November survey was
1
flown with PA-18 aircraft, January, February, and part of March flights
were made with a Cessna 185, and the remainder of March and April surveys
were made with a PA-18.
Moose condition evaluations were made during the April survey. A
body fat condition evaluation of each moose observed was made based on a
scale of (1) dead -due to natural mortality other than predation, (2)
bony -poor coat, slab-sided, hips and ribs obvious, (3) moderately
fat -fair coat, moderately rounded, hips and ribs not obvious, and (4)
fat -good coat, rounded shape, hips and ribs well-covered. Range use
evaluations during April were made to delineate areas of preferred or
critical winter range that would be inundated by construction of the
Devil Canyon and Watana dams. Classification of each area and boundaries
for each area were determined by the relative density of cumulative
moose tracks observed from early winter until April 23, 1975. The
classification categories were: (1) light use -occasional tracks with
little cratering, (2) moderate use -tracks and cratering common but not
dense, and (3) heavy use -tracks dense and cratering extensive. The
square miles of each range category were determined by overlaying a
mileage grid over a map showing the classified areas.
Harvest data were obtained from harvest report returns. Because
many hunters do not report where their animal was taken, reported
harvests for specific areas are usually less than actual harvests.
RESULTS
Moose Distributions During June, 1974.
A survey of the upper Susitna River and lower portions of major
tributaries was flown during June, 1974 to obtain spring moose dis-
tributions and to locate any areas with high densities of cows and
calves (calving areas). Results of these surveys are shown on Figure 1.
A high moose density was observed south of the MacLaren River, but no
other areas with high moose densities were observed. Few moose were
seen above 3,500 feet.
Moose Wintering Distributions, 1974-75.
Locations of moose observed during November, January, February,
March, and April surveys are shown on Figures 2 to 6, respectively. The
decrease in moose numbers observed with advancing winter was partly due
to less intensive survey procedures and partly due to poorer visibility
of moose as they move below timberline. A comparison of these maps
shows that, in most cases, moose moved from higher to lower elevations
along drainages as winter progressed. For example, moose seen near the
Susitna glaciers during November (Fig. 2) apparently moved down to
Valdez Creek by January (Fig. 3), and down to Windy Creek by February
(Fig. 4). One possible exception to this movement pattern from high to
low elevations within a drainage system was noted. The large moose
concentration along the "big bend" of the Susitna River observed during
2
November was not apparent during later surveys. It is possible that
these moose crossed the Susitna River to join wintering moose con-
concentration along the "big bend" of the Susitna River observed during
later surveys. It is possible that these moose crossed the Susitna
River to join wintering moose concentrations observed along the Oshetna
River and Sanona Creek during late winter. Heavy trailing on and along
major drainages was commonly observed. Trails criss-crossed drainages
within moose concentration areas, indicating that vegetation along both
banks was being utilized.
Moose Abundance and Composition.
Within the Susitna study area as defined for the 1974-75 winter
surveys, 2,225 moose were counted during intensive November surveys.
However, not all of the drainages were surveyed (Fig. 2). Extrapo-
lations for areas not counted can be made by multiplying the square
miles of each unsurveyed area times the moose density that was observed
in nearby similar habitat. Based on this procedure, we may have counted
2,826 moose if all of the Susitna study area were surveyed. In the
Gulkana drainage system observers saw 40 percent (28 of 70) of the moose
that were collared approximately two weeks prior to surveys. Assuming a
similar sightability of moose in the Susitna River drainages, 7~065
moose may have been in the Susitna study area. Calculated composition
ratios for the Susitna study area were 15 bulls per 100 cows and 26
calves per 100 cows.
Evaluation of Moose Winter Range, Moose Condition, and the Loss of Winter
Range by Inundation.
Observations of moose distribution through the winter indicated
that several habitat types were successively used as winter progressed.
During November surveys (Fig. 2), most moose were at or near timberline
or in riparian willow patches above timberline. A previous ground
survey (May 31, 1974) of the vegetation near timberline habitat within
the big bend of the Susitna River above the mouth of Goose Creek was the
basis for the following observations. This slope just below tree line
contains black spruce and alder as major tall shrubs and trees, dwarf
birch, alder, Salix alaxensis and Salix arbusculoides as important low
shrub species, and Ledum sp., Vaccinium vitis-idaea and Carex sp. as the
more important ground vegetation. Salix alaxensis , mainly found along
small drainages, was severely hedged with many decadent stems. A large
percentage of terminal twigs of other willow species were utilized, and
some .utilization of alder was observed. Small willow shrubs were
scattered among the more plentiful black spruce, dwarf birch, and alder
away from drainages, and many of these vlillows had been repeatedly
browsed by moose to snowline during previous winters. The usual snow-
line has apparently been at about 2 feet on flat portions of these
slopes, perhaps indicating substantial wind in this area in the winter.
Low bush cranberry is plentiful on this slope and is a potential food
source. The annual available forage on this slope appears great, but
Salix alaxensis has been over-utilized, and other willow species are at
3
least moderately-to-heavily utilized. Most moose observed below timberline
were also near riparian willow· habitat.
An increasing concentration of moose along the margins of larger,
lower elevation drainages had become apparent by January (Fig. 3). This
may have been partially due to increasing snow depths that reduced the
availability of lower-growing alpine willows. An increasing use of
vegetation growing on the steep slopes along the banks of the Susitna
River below Goose Creek was noted during January and February surveys
(Fig. 3 and Fig. 4). Many of the willow-supporting islands of the
Susitna River were examined, and it was speculated that most of the
available browse on these sites had been utilized, forcing the moose to
go elsewhere for food.
Ground examination of these river bottom willow-covered sandbars
were made during two different periods. A ground examination of a
willow bar at the mouth of the Tyone River during May 31, 1974 was the
basis for the following observations. We landed initially alongside a
willmv-covered river bar near the mouth of the Tyone River. Six to ten
foot tall balsam poplar with a low density of taller willows dominated
the vegetation in the center of the bar. Utilization of these willows
was light to moderate. The periphery of the bar consisted of a 2 to 3
foot high moderately dense stand of willows that appeared to be almost
evenly cropped (mainly moose cropping, some rabbit clipping) at the
presumed snow line. Fred Williams, sport fish biologist conducting the
sport fish studies at that time, stated that utilization of willows was
also high on the sand bars he has visited. During April, 1975 two
willm.,r-covered sandbars on the Susitna River below the MacLaren River
were examined and the willow bar near the mouth of the Tyone River was
revisited. These \villow bars were completely tracked over by moose.
Although maximum snow depths had receded by the time of these surveys,
it appeared that essentially all of the willow twigs above snowline had
been cropped. A moose calf that had starved was lying on the Tyone
River sandbar.
By late April, there were relatively few moose or moose tracks
crossing the Susitna River below the mouth of the Tyone River. The snow
had accumulated to above normal depths in the northern portion of the
Susitna study area, and most moose were observed in relatively large
concentrations. Moose range was evaluated during April and was placed
into light, moderate, or heavy use categories depending on the density
of cumulative tracking and cratering (Fig. 6). The contour intervals of
areas that would be inundated by the proposed Devil Canyon and Watana
Creek dams were superimposed on these moose range maps, and categories
of moose range that would be inundated were measured to obtain the
following results.
4
Proposed Dam
Devil Canyon
Watana
Combined
Maximum
Water Level
1450
2045
Moose Range
Category
Light
Moderate
Heavy
Light
Moderate
Heavy
Light
Moderate
Heavy
Area Indundated,
Sq. Mi.
6.8
5.6
0
0
20.2
44.0
6.8
25.8
44.0
Our data indicated that 12.4 mi.2 would be inundated by the Devil Canyon
Dam (vs 11.8 mi.2 calculated by the U.S. Corps of Engineers) and 64.2 mi.2
would be inundated by the Watana Dam (vs 67.1 mi.2 calculated by the
U.S. Corps of Engineers). It is assumed that the differences are due to
our necessarily crude methods of measuring areas. It is apparent that
the Devil Canyon Dam will have less serious consequences by inundation
of moose winter range than the Watana Dam. Examination of Figure 6 shows
that any flooding of the Susitna River above Deadman Creek will result
in the loss of heavy or moderately-used moose winter range.
Moose body condition was evaluated to compare moose in different
drainages and to see how well moose fared during the 1974-75 winter.
Samples were too small to compare moose in different drainages, so the
pooled results for the upper Susitna study area are shown below.
Percent (No.) of Moose
Area Condition Rating Adults Calves
Combined Coal Creek, Dead: o% (1) 3% ( 1)
MacLaren River, and Bony: 18% ( 21) 72% ( 26)
Clearwater Creek. Moderate: 65% (75) 25% (9)
Fat: 17% ( 20) ( 0)
5
This information shows that the wintering areas used by adult moose
during the 1974-75 winter (with above average snowfall) were adequate to
maintain them in a moderately fat condition, but moose calves became
food limited. An assessment of moose wintering on the Oshetna River
indicated that the adults were moderatly fat but snow was shallower and
browse was more available in comparison to the Clearwater Creek -
MacLaren River area.
Caribou Distributions and Trails.
Observations of caribou during the winter surveys are shown on
Figures 2 to 5. Generally, few caribou wintered in the Susitna study
area. Several hundred caribou have been observed on the Susitna River
above the Denali Highway and the adjacent higher country between Valdez
Creek and the East Fork of the Susitna River during previous November
surveys. A total of 255 were seen in this area during November 1974
(Fig. 2) but they were not seen during subsequent monthly surveys. In
addition to the caribou groups shown in Figures 2 to 5, tracks of a band
of caribou located just south of Devil Canyon during November (Fig. 2)
indicated that perhaps 50-100 caribou were in that vicinity.
The observation of well-defined, rutted caribou trails crossing the
Susitna River east of Watana Creek (Fig. 2) were of especial interest.
These trails were observed on opposite banks of the Susitna River,
indicating this is a traditional crossing area. Other trails north of
Watana Mountain led to the Susitna River but could not be found on the
opposing north bank. A substantial portion of the Nelchina caribou herd
(numbering from 8,000 to 60,000 during the last twenty years) usually
appears around the Deadman Lake -Butte Lake area during the summers,
and it is possible that these animals may frequently use the observed
crossing site of the Susitna River. No rutted trails crossing the
Susitna River were seen elsewhere during the 1974-75 surveys.
Harvest.:::; and Hunting Pressure.
Reported harvests of moose, caribou and sheep and annual numbers of
moose hunters are shown in Table 1. Since 1963, an average of 1,315
moose have been harvested annually from Unit 13 by an average of 3,666
hunters. A ratio of moose killed in the Susitna study area to moose
killed in the center of Unit 13 was derived from 1974 harvest reports;
if that ratio was constant in past harvests, the Susitna study area
would have yielded an average of 413 moose annually harvested from the
upper Susitna River drainages. Variance in hunter harvest reports over
the years does not provide all data needed to fully qualify that figure.
Estimated caribou harvests from Unit 13 based on harvest reports
indicate that an average of 5,386 caribou annually have been harvested
since 1963. The portion of this kill from the upper Susitna River
drainages has probably varied \videly over the years, but it may have
approximated one-third of the average annual harvest from Unit 13.
6
The reported harvest from the Watana Hills Dall sheep herd is
usually about 3 sheep.
Observations of Other Mammals.
A group of approximately 200 Dall sheep inhabit the range of hills
lying east of Watana Creek -Butte Creek and west of Jay Creek -Coal
Creek. These sheep are partially isolated from the larger sheep population
of the Talkeetna Mountains by low country. Although immigrations and
emigrations may occasionally be expected, in most years the Watana Hills
sheep herd is probably distinct. A portion of this sheep herd was seen
during the April survey (Fig. 6), even though no effort was made during
the surveys to fly at the higher elevations where sheep sightings would
be expected.
Wolves, wolverines, and foxes were frequently seen distributed
throughout the Susitna study area, but observations are not recorded
here.
DISCUSSION AND CONCLUSIONS
Surveys to obtain moose distributions have shown moose to generally
be at low elevations in the late winter and spring and at higher elevations
in the late fall and early winter. The proposed Susitna River dams,
therefore, may effect moose in entire drainage systems and not merely
those moose seen within or near the areas of inundation.
Those situtaions where many moose have crossed or traveled along
river corridors that will be flooded or will have fluctuating water or
ice levels are of particular concern. As an example of major river
crossings, the available information suggests that most moose seen
during early winter within the "big bend" north of the Susitna River
cross the Susitna River to join moose wintering on the lower Oshetna
River vicinity. These moose may still mostly be south of the Susitna
River during June. As another example, the dense moose concentration
seen south of the MacLaren River during June may be mainly the same
wintering moose concentration that was found during April on Clearwater
Creek. Prevention of these seasonal movements may result in a sharp
reduction in numbers of the affected moose. Ice shelves created by
fluctuating water levels in the winter or deep, wide impoundments may
act as complete or partial barriers to movements.
In addition to river crossings as part of seasonal migrations, the
criss-crossing of rivers by moose that spend a portion of the winter
near rivers is of concern. Tracks indicated that moose use vegetation
on both sides of streams, and it seems possible that prevention of moose
crossings may lower local carrying capacity by (1) isolating pockets of
vegetation where ready access is only via the frozen river and (2)
creating localized pockets of browse that are insufficient in quantity
to attract and support moose but would have contributed to the support
of those moose attracted by additional nearby browse.
7
Moose generally appeared to successively use different habitat
types during the winter. During early winter, most moose were near
timberline, but they were found increasingly at lower elevations among
riparian browse and along the steep slopes of the Susitna River by
midwinter. By late winter, the steep slopes of the Susitna River and mid-
elevations along the Susitna River, that had previously supported moose,
were infrequently used and more moose were mostly found in larger con-
centrations in willow patches on the Susitna River's major tributaries.
Following snow recession during the spring, most moose were thinly
distributed at lower elevations except for a concentration area south of
the MacLaren River. While the importance of some areas to moose may be
proportional to the extensiveness, quality, and availability of contained
browse, some areas may be of importance out of proportion to the contained
browse depending on the winter snow accumulation, slope, time of leafing
out of browse, or other factors. The relevance of this possibility is
suggested by the observed shifting concentrations of moose in various
areas of the Susitna River or its major tributaries at different time
periods.
Over 7,000 moose may have been within the study area. Natural
mortality due to predation is probably high and calf survival over the
last decade has been low. The contained moose population may be somewhat
below its optimum size.
The Susitna study area below the Denali Highway was not utilized by
substantial numbers of wintering caribou. However, a large portion of
the Nelchina caribou herd traditionally crosses the Susitna River from
its calving area near Kosina Creek to spend the summer in the Deadman
Lake -Butte Lake vicinity. A major crossing site on the Susitna River
was located just east of Watana Creek. The Susitna River appears to be
a formidable obstacle to calf caribou. Changing of conditions at this
crossing may or may not prevent the passage of adult caribou, but the
effects on calves as they attempt to follow the cows must also be
considered. Should modifications of this crossing site make the Susitna
River a barrier to caribou passage, the loss of habitat would directly
lower the potential maximum population size. Secondarily, a reduction
in recreational value of the upper Susitna River would result from the
loss of recreational caribou hunting.
The Watana Hills sheep herd lies within the Susitna study area, but
these sheep will probably not be directly affected by construction of
dams on the Susitna River. Other big game or fur bearer populations
would probably be impacted by indirect effects of increased human access
and altered numbers of prey species, but these potential impacts were
not studied and are presently unknown.
From the standpoint of recreational hunting, the Susitna study area
may be one of the most important areas in the state. Harvest data show
that the Susitna study area contributes a token sheep harvest but a
moderately large moose harvest. Most of the moose harvest from the
8
Susitna study area is from the Denali Highway -Coal Creek vicinity and
from the upper Oshetna River vicinity. Access has rapidly been increasing
in recent years, and the central portion of this area will probably
contribute to an increasing extent if past access trends continue. The
usual contribution of the Susitna study area to the annual caribou
harvest was assessed as perhaps one-third of the total. During the past
three years, most moose and caribou hunting activity within Unit 13
appeared to be on both the north and south sides of the Susitna study
area.
An indirect effect that would probably result from construction of
Susitna River dams would be increased access into the center of Game
Management Unit 13 through road construction and waterway access.
Although this has both positive and negative implications to wildlife,
the negative aspects predominate. A major increase in access would
probably require more intensive management activities with a resulting
increase in wildlife management costs. A highway corridor alongside the
Susitna River may increase the potential barrier to caribou movements.
In addition, any increased human activity near the Nelchina caribou's
calving grounds is undesirable.
In summary, moose and caribou are the key wildlife assets of the
upper Susitna River, and the major effect of dams on these ungulates is
negative. Moose may be impacted by blockage of seasonal movements
across or along river corridors due to fluctuating ice levels or deep
water impoundments and by direct loss of critical winter range through
flooding. Caribou movements may be similarly impacted by impounded
water or fluctuating ice levels, and the Nelchina caribou calving area
will probably be exposed to more human activity secondary to better
access and dam construction activities. Wildlife management costs will
necessarily increase, and the overall effect of these dams will be to
decrease numbers of moose and caribou. The effect of the Devil Canyon
Dam alone will be nild; the effect of the Watana Dam is expected to be
moderately severe. Any dam on the Susitna River that impounds water
above Deadman Creek will inundate moderately or heavily-used moose
winter range; any dam that impounds water above Watana Creek may disrupt
moose and caribou movements with potentially severe effects.
The scope of this paper does not extend to downstream wildlife or
the effects that the dam would have on those species; effects may prove
considerable.
MITIGATIVE ACTIONS
Prior to dam construction activities, detailed studies should be
conducted to more fully determine the use of this area by resident
wildlife, to gain a better understanding of the potential effects of
dams on the area's vegetation and wildlife, and to evaluate range
improvement techniques for possible use to offset loss of moose range.
Ungulate movements across drainages are largely seasonal. Where operation
of dams results in fluctuating ice levels that may impede wildlife
9
movements, changes in timing of these operations perhaps could be made
that would exchange a loss of operating efficiency for a reduced barrier
to ungulate movements. Loss of moose winter range may be partially
compensated for by well-planned, extensive range rehabilitation over a
long period of time. However, even a good and extensive range improvement
program probably won't fully mitigate any substantial losses of riparian
willow habitat.
10
REFERENCES CITED
Anon. 1954. A progress report on wildlife of the Susitna River basin.
35 pp. U.S.D.I. Fish and Wildlife Service, Juneau.
Anon. 1960. A detailed report on fish and wildlife resources affected
by the Devil Canyon Project, Alaska. 26 pp. U.S.D.I. Fish and
Wildlife Service and Bureau of Commercial Fisheries, Juneau.
11
Table 1. Harvest Data from Game Management Unit 13.
1963 1964 1965 1966, 1967 1968 1969 1970 1971 1972 1973 1974
Reported Moose Harvest, Unit 13: 1735 1607 1331 1553 1552 1512 1219 1329 1815 ':'712 618 794
Reported Moose Harvest, Center Unit 13a: 578 691 299 353 506 512 405 427 540 302 324 394
Estimated Moose Harvest from b upper Susitna River drainages 537 642 278 328 470 476 376 397 502 281 301 366
Total Hoose Hunters, Unit 13: 4163 4027 4476 3381 3585 4881 3199 2513 2770
Estimated Caribou Harvest, Unit 13: 6300 8000 7100 5500 4000 6000 7800 7247 10,131 555 810 1192
Reported Sheep Harvest, Watana Hills: 5 1 7 2 2 2 3
a Actual harvests are higher because of harvests where location of kill was not reported. The center of Unit 13 is
that portion of Unit 13 bounded by the Glenn, Richardson, Denali, and Anchorage-Fairbanks Highway.
b Estimated harvests from the upper Susitna River drainages during past years were obtained by multiplying annual
moose harvests from the center of Unit 13 times the 1974 ratio of (moose harvest from upper Susitna River drainages/
moose harvest in the center of Unit 13).
Figure 1. Moose Distributions Seen During
June 1974 Survey.
• Moose
~ Area Surveyed
~~,-.,1--· _.....-,
)0 /'
..
I r
''·.,./I U
I -l
-~-
0
02552
Q
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~
November 1974 Moose Composition Count.
Areas within broken lines were not counted.
• Moose, 0 5 Caribou, <=>Sheep
/JQ Traditional Caribou Crossing Area
I
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; 5.;
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•' ·, J ., Mo'ose concentrations during the January flight
1
Figure 3. of the Susitna Project. 1975
·-<> Hoose 0 Caribou Q Sheep
~-~::;\"?~---·\-zr-~-:-:~f
.• ~~-y~·' \Y.z~~-~-:r-7 -i-MO'NlAI
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0
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Moose concentrations during the February flight
of the Susitna Project. 1975
• Moose, 0 Caribou, 0 Sheep
·s
D
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/ 1 .. .i'--~·· \ 'I
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ouri
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rol
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i 7952 y ' Figure
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Pass'
",•)\ ..... ,,7'
I MOose concentration·s during the-':i\pril -flight, and, areas of
6. light, moderate, and heavy utilization by moose. Areas
surrounded by the broken lines are the proposed inundated
areas. 1975
I.
• Moose,
Utilization Key:
0 Caribou,
~ Light Utilization
c::z=;;·Moderate Utilization
J1l11]( Heavy Utilization
0 Sheep
: ··'):.
I (~187( 1:
-"-OENAtl RD. COMM NO \--<..J 2525 1' 12··
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