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Susitna-Watana Hydroelectric Project Document
ARLIS Uniform Cover Page
Title:
Cook Inlet beluga whale and prey study
SuWa 137
Author(s) – Personal:
Author(s) – Corporate:
Alaska Energy Authority
AEA-identified category, if specified:
Aquatic and fish resources study requests
AEA-identified series, if specified:
Series (ARLIS-assigned report number):
Susitna-Watana Hydroelectric Project document number 137
Existing numbers on document:
Published by:
[Anchorage] : Susitna-Watana Hydroelectric Project, [2012]
Date published:
5/15/12
Published for:
Date or date range of report:
Volume and/or Part numbers:
Final or Draft status, as indicated:
Document type:
Pagination:
9 p.
Related work(s):
Pages added/changed by ARLIS:
Notes:
All reports in the Susitna-Watana Hydroelectric Project Document series include an ARLIS-
produced cover page and an ARLIS-assigned number for uniformity and citability. All reports
are posted online at http://www.arlis.org/resources/susitna-watana/
Susitna-Watana Hydroelectric Project FERC #14241 Alaska Energy Authority
Cook Inlet Beluga Whale and Prey Study Request 5/15/12 Page 1
1.1. Cook Inlet Beluga Whale and Prey Study
1.2. Requester of proposed study
AEA anticipates resource agencies will request this study.
1.3. Responses to study request criteria (18 CFR 5.9(b))
1.3.1. Describe the goals and objectives of each study proposal and the information
to be obtained.
The goal of this study is to provide relevant information on Cook Inlet Beluga Whales (CIBWs)
and prey species ecology in the Susitna River and delta with respect to potential changes in the
lower river habitat that may result due to construction and operation of the Susitna-Watana
Hydroelectric Project (Project). Prey species information for eulachon will be gathered as a part
of this study while information for salmon will be coordinated with other fish studies ongoing and
proposed for the lower river. This information will be used by the Federal Energy Regulatory
Commission (FERC) during National Environmental Policy Act (NEPA) documentation and the
Project licensing process; for the National Marine Fisheries Service (NMFS) Marine Mammal
Protection Act (MMPA) and Endangered Species Act (ESA) consultations; and for the
development of avoidance, minimization, and mitigation measures if necessary.
There are seven objectives identified for this study.
1) Summarize existing information on life history, run timing, abundance, distribution, and
habitat of beluga whale prey species in the Susitna River and in other Cook Inlet
tributaries used by beluga whales.
2) Summarize existing information on the temporal and spatial distribution of beluga
whales in Cook Inlet, the Susitna River delta, and the Susitna River relative to the
availability of eulachon, Chinook, sockeye, chum, and coho salmon.
3) Provide a factual basis to Iinitiate consultation with the NMFS for Marine Mammal
Protection Act (MMPA) and Endangered Species Act (ESA) permitting and
requirements for the Susitna Watana study program.
4) Document the current presence and the upstream extent of Cook Inlet beluga whales
within the Susitna River delta.
5) Describe the current densities of adult eulachon and their spawning habitat in the
lower Susitna River.
6) Evaluate the relationship between potential hydropower-related changes in the lower
river and in-river movements of beluga whales.
7) Evaluate the relationship between potential hydropower-related changes in the lower
river and eulachon spawning habitat.
1.3.2. If applicable, explain the relevant resource management goals of the agencies
and/or Alaska Native entities with jurisdiction over the resource to be studied.
The NMFS is required to promote the conservation of all marine mammal species, particularly if
they are designated as depleted under the MMPA or listed as threatened or endangered under
the ESA. CIBWs are protected both as a depleted stock under the MMPA and as endangered
under the ESA. Traditionally, CIBW were a subsistence resource for Alaska Natives, however,
the subsistence harvest was limited in 1999 due to sharp declines in CIBW abundance
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Cook Inlet Beluga Whale and Prey Study Request 5/15/12 Page 2
throughout the 1990’s. The CIBW population has been below the harvestable level of 350 since
2009 (Allen and Angliss 2012).
1.3.3. If the requester is not a resource agency, explain any relevant public interest
considerations in regard to the proposed study.
Fisheries and marine mammal resources are owned by the State of Alaska, and the Project
could potentially affect these public interest resources by affecting flows or habitat in the Susitna
River and delta.
1.3.4. Describe existing information concerning the subject of the study proposal,
and the need for additional information.
The majority of information regarding the current CIBW population abundance and distribution
comes from annual, aerial surveys conducted by the NMFS-NMML (National Marine Mammal
Laboratory) during June and August (NMFS 2008, 2012). Additionally, aerial surveys for beluga
whales were completed in 1982 and 1983 as part of the original licensing effort (Harza-Ebasco
1985). These surveys all recognize the summer aggregations of belugas in the Susitna River
delta (NMFS 2008). Additionally, satellite tags were applied to 15 whales from 1999-2002 which
further confirmed the use of the delta by CIBWs (Hobbs et al. 2005, 2008). Finally, land-based
and boat-based surveys directed at movement and residency patterns have been conducted in
the Susitna Flats and adjacent areas (Funk et al. 2005; Prevel-Ramos et al. 2006, Markowitz
and McGuire 2007, Markowitz et al. 2007, Nemeth et al. 2007).These studies have provided a
photo-identification catalog for CIBW. Photographs of marked individuals have been used to
characterize distribution and habitat use by individuals and groups of whales (McGuire et al.
2008, McGuire and Kaplan 2009, McGuire et al. 2009, 2011a,b).
Whale movement and habitat use studies using satellite telemetry and hydrodynamic modeling
indicate that beluga distributions are controlled not only by water temperature and ice coverage,
but also by the seasonal flow patterns of various rivers. This suggests that availability of salmon
and other fish (i.e. eulachon) in river mouths influence beluga movements (Ezer 2011). Beluga
whales use the Susitna River delta throughout the majority of the open water season (late-April
through September) (NMFS 2008). The spring timing is coincident with the spawning migrations
of eulachon and Pacific salmon into the river. Availability of prey species was one of the primary
constituent elements used to designate critical habitat in 2011. Critical habitat for CIBW includes
the Susitna River delta and extends upriver approximately 8.5 NM from mean lower low water
(MLLW).
Preliminary data from studies of eulachon in the 1980s indicate that their spawning
requirements are broad and that most spawning occurs below RM 28, but some spawning
activity can be found as far upstream as RM 50 (HDR Alaska, Inc. 2011). Eulachon may be
commercially harvested in the salt waters of the Upper Cook Inlet between the Chuit and Little
Susitna Rivers from May 1 to June 30 using a hand operated dip net; the harvest is not allowed
to exceed 100 tons (ADF&G 2012). Harvest rates over the period 2006 to 2010 averaged 55
tons (Shields 2010). However, recent data on eulachon densities and distribution in the Susitna
River delta is sparse.
ADF&G has conducted ongoing salmon escapement studies in the Susitna River drainage basin
including: helicopter and foot surveys to index counts of Chinook salmon; coho salmon
escapement surveys; and coho and chum salmon telemetry studies. The Susitna River Salmon
Run Apportionment Study describes additional capture and tagging efforts on the Susitna River
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Cook Inlet Beluga Whale and Prey Study Request 5/15/12 Page 3
that will augment ADF&Gs telemetry studies of chum and coho salmon and will also mark
Chinook and pink salmon.
1.3.5. Explain any nexus between project operations and effects (direct, indirect,
and/or cumulative) on the resource to be studied, and how the study results
would inform the development of license requirements.
Cook Inlet beluga whales could be impacted by potential Project-induced changes to the
abundance, productivity, and run timing of important Susitna River prey species: eulachon, and
Chinook, sockeye, chum, and coho salmon. These Project- related effects could occur due to
changes in discharge if the changes prevented, impaired or delayed beluga whale access to
delta or river habitats that support known prey species, including eulachon and salmon. In
addition, potential effects of flow that, in turn, affect abundance, densities, and/or run timing of
these prey species also have the potential to affect CIBW. The CIBW Distinct Population
Segment is protected by the NMFS as both a depleted stock under the MMPA, and as an
endangered species under the ESA. In addition, large portions of upper Cook Inlet, including
the Susitna River delta, have been designated as critical habitat for the CIBW. Eulachon and
adult Chinook, sockeye, chum and coho salmon are identified as primary constituent elements
of CIBW critical habitat in Cook Inlet.
This study addresses the following issue identified in the PAD (AEA 2011):
• F10: Potential impacts to the Endangered Cook Inlet beluga whale.
Studies will be conducted to evaluate the impacts of Project-related changes to sediment
transport and delivery, stream temperature, water quality, stream flow, and ice processes on
salmon habitat, productivity, abundance and run timing. This study will synthesize the available
information on eulachon to understand how the Project may affect the species in the Susitna
River. The temporal and spatial distribution of beluga whales within the Susitna River will be
used to inform the assessment area of 2013–2014 studies.
1.3.6. Explain how any proposed study methodology (including any preferred data
collection and analysis techniques, or objectively quantified information, and a
schedule including appropriate field season(s) and the duration) is consistent
with generally accepted practice in the scientific community or, as appropriate,
considers relevant tribal values and knowledge.
Objective 1: Summarize existing information on life history, run timing, abundance,
distribution, and habitat of beluga whale prey species in the Susitna River and in other
Cook Inlet tributaries used by beluga whales.
This task will involve identifying, compiling and reviewing existing literature to prepare a white
paper describing the life history and habitat requirements of eulachon in glacial river systems.
Available data on the Upper Cook Inlet eulachon, Chinook, chum, coho, and sockeye salmon
fisheries and Susitna River eulachon, Chinook, chum. coho, and sockeye salmon escapement,
life history and distribution (below RM 50) will be synthesized to identify key habitat
requirements (e.g. stream temperatures, stream flows) for critical periods including spawning
(late April through June) and outmigration (June through August). Finally, we will identify any
remaining data gaps and evaluate the potential for Project impacts on eulachon and salmon
habitat to determine future study needs.
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Cook Inlet Beluga Whale and Prey Study Request 5/15/12 Page 4
Objective 2: Summarize existing information on the temporal and spatial distribution of
beluga whales in Cook Inlet, the Susitna River delta, and the Susitna River relative to the
availability of eulachon, Chinook, sockeye, chum, and coho salmon.
This task involves compiling existing temporal and spatial distribution data for Cook Inlet beluga
whales in and around the Susitna River and Susitna River delta, especially during spawning
periods of Susitna River eulachon, Chinook, sockeye, chum and coho salmon. Data will be
requested from the Port of Anchorage, Joint Base Elmendorf Richardson, Cook Inlet Region
Incorporated, and the NMFS. However, these entities may not be able to provide data and/or
may not be able to meet the desired project schedule. Spatial data products will be delivered in
the two-dimensional Alaska Albers Conical Equal Area projection, and North American Datum of
1983 (NAD 83) horizontal datum consistent with ADNR standards. Naming conventions of files
and data fields, spatial resolution, and metadata descriptions will comply with the ADNR
standards established for the Susitna-Watana Hydroelectric Project. This information will be
used to identify potential Project-induced impacts to CIBWs and critical habitat and to identify
critical data gaps to determine future study needs..
Objective 3: Initiate consultation with the NMFS for Marine Mammal Protection Act
(MMPA) and Endangered Species Act (ESA) permitting and requirements for the Susitna
Watana study program.
In association with the NMFS, determine MMPA and ESA permit needs for the environmental
studies program. Permits are required for all studies conducted within CIBW-designated critical
habitat (up to RM 9), all studies that may have flights to transport personnel and/or equipment
over open water (up to RM 9) and for CIBW surveys throughout Type 1 critical habitat planned
for spring 2013. The schedule and scope of the permits may be altered during agency
consultation. Additionally, we will develop a “No Impact” protocol to be implemented by all
projects with 2012 field studies since the project does not have MMPA or ESA permits for 2012.
Objective 4: Document the current presence and upstream extent of Cook Inlet beluga
whales in the Susitna River delta.
We will examine a combination of methods to monitor the presence, timing, and upstream
extent of CIBW in the Susitna River. CIBWs swim into tributaries of upper Cook Inlet, including
the Susitna River and adjacent Little Susitna River to feed on presumably adult and juvenile
salmon and eulachon. Historic records indicate that beluga whales have been observed in the
eastern channel of the Susitna River as far as 30-40 miles upriver, but are most commonly
found within the first 5 miles (Funk et al. 2005). Potential monitoring methods will include a
combination of systematic and opportunistic surveys. For example, visual observations from
boats by the fisheries crews based in field camps at Flathorn and at the confluence of the
Yentna River, dedicated CIBW aerial surveys, and land-based remote camera systems. The
timing and methods used for monitoring will be examined during the completion of the 2012
Study Plan (i.e., data synthesis for CIBWs and prey species). We will also investigate the
anticipated timing of Project effects on critical habitat.
Objective 5: Describe the current densities of adult eulachon and their spawning habitat
in the lower Susitna River.
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Cook Inlet Beluga Whale and Prey Study Request 5/15/12 Page 5
Because eulachon are considered broadcast spawners, they may be migrating, holding and
spawning at depths that pose challenges for fish collection. Additionally, we are interested in a
sampling protocol for eulachon that will pose the least amount of physical disturbance to
potential spawning habitat. Consequently, our primary approach to collecting estimates of
eulachon density will be by mobile (boat) surveys with a combination side- and down-looking
split-beam sonar, which has been successfully used to characterize the seasonal distribution
and density of forage fish in the Susitna River delta (Nemeth et al. 2007). Sonar transects will
be established throughout the lower river. Sampling locations will be spread throughout the
lower river and will include areas that have previously been documented as eulachon spawning
habitat and/or areas with similar physical characteristics to these known spawning locations as
well as areas where spawning has not previously been documented Transects will be located to
ensure collection of data across representative channel and/or habitat types (e.g. mainstem
riffles, side channel glides). This is the preferred method for density estimation as it will require
minimal handling and disruption to spawning eulachon, and will be able to cover large areas on
a relatively frequent basis.
As there will be other fish species staging in the lower Susitna River at the time of the eulachon
spawning migration (e.g., salmon smolts), it will be necessary to obtain species verification for
sonar-derived fish densities. Thus, a fish capture survey will be conducted in one location along
each sonar transect. Sampling for eulachon will be conducted using a variety of methods
including seining, gill netting, trawling, and/or boat electrofishing. Total catch by species, area
sampled, and measurement of effort (e.g. set times for nets, power, time, and distance for
electrofishing) will be recorded for each sampling location. If multiple species are collected, size
measurements will be obtained to help differentiate acoustic targets.
During sonar and fish sampling the crew will make observations of any marine mammals
present, particularly CIBW. All CIBW observations will be documented and locations marked
with a GPS waypoint. This information will be incorporated into the data collection effort for
Objective 1.
Objective 6: Evaluate the relationship between potential hydropower-related changes in
the lower river and in-river movements of beluga whales.
Data collected during Objectives 2 and 4 (past and present distribution and movements of CIBW
in the Susitna River delta) will be combined with data from In-stream Flow, Ice Processes and
Geomorphology studies to evaluate the potential effects of the project. The method(s) used to
evaluate the significance of potential impacts to CIBW will be dependent upon the nature and
degree of the changes in flow and channel morphology which may affect in-river movements of
CIBWs.
Objective 7: Evaluate the relationship between potential hydropower-related changes in
the lower river and eulachon spawning habitat.
The initial step in addressing this objective is to characterize eulachon spawning habitats in the
lower Susitna River (Objective 1). Historical data on eulachon spawning habitat (e.g., 1980s
data) will be synthesized through the completion of the 2012 Study Plan. In 2013-14, habitat
data will be collected concurrent with fish sampling for species verification and will be
supplemented with dedicated habitat surveys, as appropriate. Spawning habitats will be
delineated to the finest scale possible, i.e. channel location and mesohabitat type, with
upstream and downstream borders marked by GPS waypoints. Several habitat parameters will
be collected for all delineated habitats including average water depth, average wetted width,
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Cook Inlet Beluga Whale and Prey Study Request 5/15/12 Page 6
water temperature, velocity, tide elevation, and substrate composition. These habitat data will
be used as inputs to any flow-habitat models that may need to be developed to evaluate
potential changes in eulachon spawning habitat as a function of any Project-related changes in
flow and surface water elevation.
1.3.7. Describe considerations of level of effort and cost, as applicable, and why any
proposed alternative studies would not be sufficient to meet the stated
information needs.
The exact schedule, staff assignments, and costs will be detailed as the 2013–2014 Study Plan
develops. The level of effort and cost can change significantly based on the final study plans.
For instance, aerial surveys for CIBW distribution in the Susitna River delta will require 2-4
individuals for observation and data analysis. Depending on the area covered during each
survey, the type of data collected (i.e. visual observation versus photographic data) and
subsequent data analyses, systematic surveys can cost between $5,000 and $20,000 per
survey. Surveys could occur on a weekly or monthly basis year-round resulting in a cost of
$200,000 to $1,000,000 per year depending on survey design. There is similar uncertainty in
estimating costs for eulachon studies. Ideally, multiple survey crews of four people would
sample for 8 hours each day (4 hours before and after high tide). The timing of surveys should
coincide with documented eulachon runs in May and June. If similar effort to the studies
completed in the 1980’s is expected (i.e. 4 surveys per week), then costs could be in excess of
$150,000 per year for labor costs plus the cost for data analyses. Costs for equipment will also
be drastically different depending on methods that will be permitted (i.e. boat electroshocking,
side-scan sonar systems, and/or netting equipment). Lower levels of effort will reduce the cost
proportionally with effort.
1.3.8. Literature Cited
ADF&G (Alaska Department of Fish and Game). 2012. 2010 Upper Cook Inlet Commercial
Smelt (Hooligan) and Herring Fishing Seasons. Alaska Department of Fish and Game
Division of Commercial Fisheries News Release. April 6, 2010. Pat Shields Assistant Area
Management Biologist, Soldotna, Alaska. Available online at:
http://www.adfg.alaska.gov/static/fishing/PDFs/commercial/2010_uci_herringsmelt_seasons.
pdf. Accessed on January 18, 2012.
AEA (Alaska Energy Authority). 2011. Pre-Application Document: Susitna-Watana
Hydroelectric Project FERC Project No. 14241. December 2011. Prepared for the Federal
Energy Regulatory Commission by the Alaska Energy Authority, Anchorage, Alaska.
Allen, B.M., and R. P. Angliss. 2011. Alaska marine mammal stock assessments, 2010. U.S.
Department of Commerce, NOAA Tech. Memo. NMFSAFSC-223, 292 p.
Ezer, T. 2011. Using inundation modeling and remote sensing data to study hydrodynamic and
environmental impacts on the survival of Cook Inlet’s Beluga whales. Final Report for the
period: 1-October, 2010 to 30-Deptember, 2011. Project RFQ#
NOAA_NMFS_AKR_10_0802_2G. Prepared for National Oceanic & Atmospheric
Administration by Center for Coastal Physical Oceanography, Old Dominion University
Norfolk, VA. 14 pp.
Flannery, B. F., J. K. Wenburg, C. J. Lewis, B. L. Norcross, and R. E. Spangler. 2009.
Genetic population structure of Alaska eulachon. U.S. Fish and Wildlife Service, Alaska
Fisheries Technical Report 106, Anchorage.
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Cook Inlet Beluga Whale and Prey Study Request 5/15/12 Page 7
Funk, D.W., T.M. Markowitz, and R. Rodrigues, eds. 2005. Baseline Studies of Beluga Whale
Habitat Use in Knik Arm, Upper Cook Inlet, Alaska, July 2004-July 2005. Rep. from LGL
Alaska Research Associates, Inc., Anchorage, AK, in association with HDR Alaska, Inc.,
Anchorage, AK, for the Knik Arm Bridge and Toll Authority, Anchorage, AK, the Department
of Transportation and Public Facilities, Anchorage, AK, and the Federal Highway
Administration, Juneau, Alaska.
Goetz, K.T., Rugh, D.J., Read, A.J., and Hobbs, R.C. 2007. Habitat use in a marine
ecosystem: beluga whales Delphinapterus leucas in Cook Inlet, Alaska. Marine Ecology
Progress Series 330:247-256.
Harza-Ebasco (Harza-Ebasco Susitna Joint Venture). 1985. Fish Wildlife and botanical
resources. Exhibit E, Volume 9. Prepared for the Alaska Power Authority by the Harza-
Ebasco Susitna Joint Venture, Anchorage, Alaska.
HDR Alaska, Inc. 2011. Susitna-Watana Hydroelectric Project, Railbelt Large Hydro: Aquatic
Resources Data Gap Analysis. Draft Report, July 20, 2011. Prepared for The Alaska
Energy Authority by HDR Alaska, Inc., Anchorage, Alaska. 72 pp.
Hobbs, R. C., J. M. Waite, and D. J. Rugh. 2000. Beluga, Delphinapterus leucas, group
sizes in Cook Inlet, Alaska, based on observer counts and aerial video. Marine Fisheries
Review 62(3):46-59.
Hobbs, R.C., Shelden, K.E., Rugh, D.J., and Norman, S.A. 2008. 2008 status review and
extinction risk assessment of Cook Inlet belugas (Delphinapterus leucas). AFSC Processed
Report 2008-02, 116 p. Alaska Fish. Sci. Cent., NOAA, Natl. Mar. Fish. Serv., 7600 Sand
Point Way NE, Seattle, WA 98115.
Hobbs, R.C., Laidre, K.L., Vos, D.J., Mahoney, B.A., and Eagleton, M. 2005. Movements and
area use of belugas, Delphinapterus leucas, in a Subarctic Alaskan estuary. Arctic
58(4):331-340.
Huntington, H.P. 2000. Traditional Knowledge of the Ecology of Belugas, Delphinapterus
leucas, in Cook Inlet, Alaska. Marine Fisheries Review. 134-140.
LGL Alaska Research Associates, Inc. 2009. Photo-identification of beluga whales in Upper
Cook Inlet, Alaska: Mark analysis, mark-resight estimates, and color analysis from
photographs taken in 2008. Report prepared by LGL Alaska Research Associates, Inc.,
Anchorage, AK, for National Fish and Wildlife Foundation, Chevron, and ConocoPhillips
Alaska, Inc. 99 p. + Appendices.
Markowitz, T.M., and McGuire, T.L., eds. 2007. Temporal-spatial distribution, movements and
behavior of beluga whales near the Port of Anchorage, Alaska. Report from LGL Alaska
Research Associates, Inc., Anchorage, AK, for Integrated Concepts and Research
Corporation and the U.S. Department of Transportation Maritime Administration.
Markowitz, T.M., McGuire, T.L., and Savarese, D.M. 2007. Monitoring beluga whale
(Delphinapterus leucas) distribution and movements in Turnagain Arm along the Seward
Highway. Final Report. Report from LGL Alaska Research Associates, Inc., Anchorage,
AK, for HDR and the Alaska Department of Transportation and Public Facilities.
McGuire, T.L., and Kaplan, C.C. 2009. Photo-identification of beluga whales in Upper Cook
Inlet, Alaska. Final report of field activities in 2008. Report prepared by LGL Alaska
Research Associates, Inc., Anchorage, AK, for National Fish and Wildlife Foundation,
Chevron, and ConocoPhillips Alaska, Inc. 28 p. + Appendices.
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McGuire, T.L., M.K. Blees, and M.L. Bourdon. 2011a. The development of a catalog of left-side
digital images of individually-identified Cook Inlet beluga whales Delphinapterus leucas.
North Pacific Research Board Final Report 910, 96 p.
McGuire, T.L., Blees, M.K., and Bourdon, M.L. 2011b. Photo-identification of beluga whales in
Upper Cook Inlet, Alaska. Final report of field activities and belugas resighted in 2009.
Report prepared by LGL Alaska Research Associates, Inc., Anchorage, AK for National Fish
and Wildlife Foundation, Chevron, and ConocoPhillips Alaska, Inc. 53 p. + Appendices.
McGuire, T.L., Kaplan, C.C., and Blees, M.K. 2009. Photo-identification of beluga whales in
Upper Cook Inlet, Alaska. Final report of belugas resighted in 2008. Report prepared by
LGL Alaska Research Associates, Inc., Anchorage, AK, for National Fish and Wildlife
Foundation, Chevron, and ConocoPhillips Alaska, Inc. 42 p. + Appendices.
McGuire, T.L., Kaplan, C.C., Blees, M.K., and Link, M.R. 2008. Photo-identification of beluga
whales in Upper Cook Inlet, Alaska. 2007 annual report. Report prepared by LGL Alaska
Research Associates, Inc., Anchorage, AK, for Chevron, National Fish and Wildlife
Foundation, and ConocoPhillips Alaska, Inc. 52 p. + Appendices.
Moore, S.E., K.E.W. Shelden, L.K. Litzky, B.A. Mahoney and D.J. Rugh. 2000. “Beluga
whale, (Delphinapterus leucas), habitat associations in Cook Inlet, Alaska.” Marine Fisheries
Review 62(3):60–80.
Nemeth, M.J., Kaplan, C.C., Prevel-Ramos, A.M., Wade, G.D., Savarese, D.M., and Lyons,
C.D. 2007. Baseline studies of marine fish and mammals in Upper Cook Inlet, April through
October 2006. Final report prepared by LGL Alaska Research Associates, Inc., Anchorage,
AK for DRven Corporation, Anchorage, AK.
NMFS. 2008. Endangered and threatened species: endangered status of the Cook Inlet beluga
whale. Federal Register 73(205):62919-62930.
Nemeth, M. J., C. C. Kaplan, A. P. Ramos, G. D. Wade, D. M. Savarese, and C. D. Lyons.
2007. Baseline studies of marine fish and mammals in Upper Cook Inlet, April through
October 2006. Final report prepared by LGL Alaska Research Associates, Inc., Anchorage,
Alaska for DRven Corporation, Anchorage, Alaska.
NMFS (National Marine Fisheries Service). 2008. Conservation Plan for the Cook Inlet beluga
whale (Delphinapterus leucas). National Marine Fisheries Service, Juneau, Alaska.
NMFS (National Marine Fisheries Service). 2012. Cook Inlet Beluga Whales. National
Oceanic and Atmospheric Administration, National Marine Fisheries Service, Alaska
Regional Office. Available online at:
http://www.fakr.noaa.gov/protectedresources/whales/beluga.htm Accessed January 14,
2012.
Prevel-Ramos, A.M., Markowitz, T.M., Funk, D.W., and Link, M.R. 2006. Monitoring beluga
whales at the Port of Anchorage: Pre-expansion observations, August-November, 2005.
Report from LGL Alaska Research Associates, Inc., Anchorage, AK, for Integrated Concepts
and Research Corporation, the Port of Anchorage, and the U.S. Department of
Transportation Maritime Administration.
Rugh, D.J., Shelden, K.E.W., and Mahoney, B.A. 2000. Distribution of belugas, Delphinapterus
leucas, in Cook Inlet, Alaska, during June/July 1993-2000. Marine Fisheries Review
62(3):6-21.
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Rugh, D.J., Goetz, K.T., Sims, C.L., Shelden, K.W., Shpak, O.V., Mahoney, B.A., and Smith,
B.K. 2006. Aerial surveys of belugas in Cook Inlet, Alaska, June 2006.
http://www.fakr.noaa.gov/protectedresources/whales/beluga/survey/june2006.pdf
Rugh, D.J., Shelden, K.E.W., Sims, C.L., Mahoney, B.A., Smith, B.K., Litzky, L.K., and Hobbs,
R.C. 2005. Aerial surveys of belugas in Cook Inlet, Alaska, June 2001, 2002, 2003, and
2004. NOAA Technical Memorandum NMFS-AFSC-149.
Rugh, D.J., Mahoney, B.A., and Smith, B.K. 2004. Aerial surveys of beluga whales in Cook
Inlet, Alaska, between June 2001 and June 2002. NOAA Technical Memorandum NMFS-
AFSC-145.
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